Environment: milieu / climate zone / depth range / distribution range
экология
; пресноводный; солоноватоводный донно-пелагический; анадромный (Ref. 51243). Subtropical; 10°C - 20°C (Ref. 2059); 64°N - 35°N, 12°E - 53°E
Eurasia: Caspian, Black, Marmara and Baltic Sea basins (Sweden and Finland north to 63°N), North Sea basin from Elbe to Ems drainages. In Anatolia: in Marmara basin, south to Great Menderes and Lake Egridir and east to Kızılırmak drainage. Absent between Kızılırmak and Çoruh drainages. Extirpated in Crimea. Introduced in Rhine.
Size / Вес / Возраст
Maturity: Lm ?  range ? - ? cm
Max length : 50.0 cm TL самец/пол неопределен; (Ref. 556); common length : 20.0 cm TL самец/пол неопределен; (Ref. 556); наибольший вес (опубликованные данные): 1.4 kg (Ref. 35541); наибольший возраст (опубликованны данные): 15 годы (Ref. 59043)
Краткое описание
определительные ключи | морфология | морфометрия
колючие лучи спинного плавника (общее число) : 3; членистые (мягкие) лучи спинного плавника (общее число) : 8 - 9; колючие лучи анального плавника: 3; членистые (мягкие) лучи анального плавника: 16 - 22. Diagnosed from its congeners in Europe by the following characters: anal fin with 16-21½ branched rays; back keeled behind dorsal base; in spawning season, males blackish brown with orange cheek and sometimes belly (Ref. 59043). Caudal fin with 19 soft rays. Scalar formula: 49-64, 6-10 (Ref. 40476).
Inhabits brackish estuaries, large to medium rivers and some large subalpine lakes. Sedentary populations inhabit even in small rivers or barbel zone. Feeds mainly on small molluscs and insect larvae. Breeds in riffles in shallow, fast-flowing streams and rivers on gravel. Semi-anadromous populations forage in freshened parts of sea and migrate for long distances to spawn. Lacustrine populations move to fast-flowing tributaries. Spent adults return to foraging habitats (Ref. 59043). Threatened from its range due to water obstruction (Ref. 26100).
Produces 15,000-20,000 eggs at the age of 6-7 years (Ref. 12259). Spawns for several years. Semi-anadromous populations undertake long distance spawning migrations while lacustrine populations migrate to fast-flowing tributaries. In Dniepr and Kuban, migration starts in September, decreases in winter and resumes in March-May (Ref. 59043).
Kottelat, M. and J. Freyhof, 2007. Handbook of European freshwater fishes. Publications Kottelat, Cornol and Freyhof, Berlin. 646 pp. (Ref. 59043)
Статус Красного Списка МСОП (Ref. 130435)
Угроза для людей
Harmless
Использование человеком
рыболовство: не имеет хозяйственного значения; аквакультура (рыбоводство): коммерческий; объект спортивного рыболовства: да
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Estimates based on models
Phylogenetic diversity index (Ref.
82804): PD
50 = 0.5625 [Uniqueness, from 0.5 = low to 2.0 = high].
Bayesian length-weight: a=0.00724 (0.00660 - 0.00795), b=3.14 (3.11 - 3.17), in cm total length, based on LWR estimates for this species (Ref.
93245).
Trophic level (Ref.
69278): 3.3 ±0.1 se; based on diet studies.
устойчивость к внешним воздействиям (Ref.
120179): низкий, минимальное время удвоения популяции 4.5-14 лет (K=0.22; tm=3; tmax=15).
Fishing Vulnerability (Ref.
59153): Moderate vulnerability (37 of 100).