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Raja clavata Linnaeus, 1758

Thornback ray
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Raja clavata   AquaMaps   Data sources: GBIF OBIS
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Image of Raja clavata (Thornback ray)
Raja clavata
Picture by Stergiou, K.I.

Classification / Names आम नाम | उपशब्द | Catalog of Fishes(वर्ग, प्रजाति) | ITIS | CoL | WoRMS | Cloffa

इलाज़मौबरांकी (शारक और रेज़) (sharks and rays) > Rajiformes (Skates and rays) > Rajidae (Skates)
Etymology: Raja: Latin, raja, -ae = a sting ray (Raja sp.) (Ref. 45335).
More on author: Linnaeus.

Environment: milieu / climate zone / depth range / distribution range पारिस्थितिकी

समुद्री ड़िमरसल; गहराई सीमा 5 - 1020 m (Ref. 114953). Subtropical; 70°N - 38°S, 25°W - 50°E

वितरण देश | ऐफ ऐ ओ क्षेत्र | Ecosystems | संयोग | Point map | भूमिका | Faunafri

Eastern Atlantic and Southwest Indian Ocean: Iceland to Madagascar, including the Mediterranean Sea.

Length at first maturity / आकार / वज़न / Age

Maturity: Lm 76.6, range 47 - 87.5 cm
Max length : 105 cm TL पुल्लिंग / अलिंग; (Ref. 53748); 139.0 cm TL (female); common length : 85.0 cm TL पुल्लिंग / अलिंग; (Ref. 9987); अधिकतम प्रकाशित वज़न: 18.0 kg (Ref. 35388); अधिकतम सूचित उम्र: 15 वर्षो (Ref. 41305)

Short description पहचान कुंजी | आकृति विज्ञान | मौरफोमैटरिक्स

पृष्ठीय रीढ़ (सम्पूर्ण) : 0; गुदा कांटा: 0. Sub-rhomboid in shape with pointed wing-tips; disc-width 1,25 to 1,36 times in its length, its length 1,70 to 1,83 times in total length. Short rostrum, rounded at his extremity; pectoral fins with clear angles on lateral side; triangular pelvic fins (Ref. 39246). Dorsally prickly; large females also prickly throughout their ventral surface; juveniles and large males prickly along the margins of their discs and the underside of their snout. 30-50 thorns form a median row from the nape to the first dorsal fin; additional large 'buckler' thorns with swollen bases scattered on upper surface of disc in adults (Ref. 3167). Coloration is highly variable, with the dorsal surface having all shades of brown with dark and light spots and blotches; ventral surface white (Ref. 3167, 58137).

जीवविज्ञान     शब्द संग्रह (उदाहरणार्थ epibenthic)

Inhabits shelf and upper slope waters. Reported depth ranges varies, from 10-300 m (Ref. 4426, 82319, 88187), and from 300-577 m in the eastern Ionian Sea (Ref. 56504); most common in coastal waters between 10-60 m depth (Ref. 58137). Tolerates low salinities (Ref. 88171). Found on mud, sand and gravel bottoms, rarely on rougher bottoms (Ref. 6808, 58137). Nocturnal species. Feeds on all kinds of bottom animals, preferably crustaceans (Ref. 3167) and fishes (Ref. 114953). Undertakes migrations with mean distances of 54-117 km per month; shows a clear annual migration cycle (Ref. 89017, 89018), moves from deeper offshore waters (10-30 m) in autumn and winter to shallower areas (<10 m) in spring (Ref. 82319). Young are non-migratory, inhabiting inshore nursery grounds (Ref. 89019, 89020, 89022); in the Bay of Douarnenez (France) they are found to remain in shallow waters for at least 2 years (Ref. 89023). Maturity age 7-8 years (Ref. 114953). Oviparous. Young may tend to follow large objects, such as their mother (Ref. 205). Detects weak electric fields generated by other organisms (prey detection and predator avoidance) but may also generate its own weak electric fields (Ref. 10311, 88171). Utilized fresh and frozen (Ref. 9987).

Life cycle and mating behavior Maturities | पुनरुत्पत्ति | Spawnings | Egg(s) | Fecundities | लार्वा

Oviparous. Distinct pairing with embrace. Polyandrous species (Ref. 89024). Paired eggs are laid and deposited on shallow sand, mud, pebble or gravel bottoms (Ref. 205, 89025). Up to 170 egg cases can be laid by a single female in a year (Ref. 3167), but average fecundity is much lower (around 48-74 eggs) (Ref. 3603, 31302). In northwestern Europe, egg cases are laid during spring (Ref. 3167) and in the Mediterranean during winter and spring (Ref. 3167). Egg cases are oblong capsules with stiff pointed horns at the corners, each containing one embryo (Ref. 205). Capsules are 5.0-9.0 cm long without the horns (Ref. 41250, 88187) and 3.4-6.8 cm wide (Ref. 41250). Egg cases are anchored with an adhesive film (Ref. 82399). Embryos feed solely on yolk (Ref. 50449). Egg cases hatch after about 4-5 months and pups are about 11-13 cm TL (Ref. 88864). Mating season from February to September, peaking in June (Ref. 74501). Adults observed to form same-sex aggregations during the mating season with females moving to shallower inshore waters approximately a month before the males (Ref 3603, 58137, 74501). Mating does not occur in the Baltic Sea (Ref. 82311).

Main reference Upload your references | संदर्भ | संयोजक : McEachran, John | सहयोगीयो

McEachran, J.D. and K.A. Dunn, 1998. Phylogenetic analysis of skates, a morphologically conservative clade of elasmobranchs (Chondrichthyes: Rajidae). Copeia 1998(2):271-290. (Ref. 27314)

IUCN Red List Status (Ref. 130435)

  Near Threatened (NT) ; Date assessed: 03 October 2005

CITES

Not Evaluated

CMS (Ref. 116361)

Not Evaluated

Threat to humans

  Harmless





Human uses

मात्स्यिकी: व्यापारिक; आखेट मत्स्य: हां
FAO - मात्स्यिकी: landings; Publication: search | FishSource | Sea Around Us

अधिक जानकारी

Life cycle
पुनरुत्पत्ति
Maturities
Fecundities
Spawnings
Spawning aggregations
Egg(s)
Egg developments
लार्वा
लारवल गतिकी
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Visual pigment(s)
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Diseases / Parasites
Toxicities (LC50s)
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References
संदर्भ

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इंटरनेट स्रोत

AFORO (otoliths) | Aquatic Commons | BHL | Cloffa | BOLDSystems | Websites from users | Check FishWatcher | CISTI | Catalog of Fishes: वर्ग, प्रजाति | DiscoverLife | DORIS | ECOTOX | FAO - मात्स्यिकी: landings; Publication: search | Faunafri | Fishipedia | Fishtrace | GenBank: genome, nucleotide | GloBI | Google Books | Google Scholar | Google | IGFA World Record | MitoFish | राष्ट्रीय आंकड़ासंचय | Otolith Atlas of Taiwan Fishes | सार्वजनिक लजीवालय | PubMed | Reef Life Survey | Socotra Atlas | Tree of Life | Wikipedia: Go, खोज | World Records Freshwater Fishing | Zoobank | Zoological Record

Estimates based on models

Preferred temperature (Ref. 123201): 7.1 - 15.8, mean 10.7 °C (based on 474 cells).
Phylogenetic diversity index (Ref. 82804):  PD50 = 0.5000   [Uniqueness, from 0.5 = low to 2.0 = high].
Bayesian length-weight: a=0.00200 (0.00175 - 0.00228), b=3.26 (3.23 - 3.29), in cm total length, based on LWR estimates for this species (Ref. 93245).
Trophic level (Ref. 69278):  3.8   ±0.2 se; based on diet studies.
Generation time: 7.8 (5.4 - 10.4) years. Estimated as median ln(3)/K based on 16 growth studies.
लौटाव (Ref. 120179):  निम्न, न्यूनतम जनसंख्या दुगनी होने का समय 4.5 - 14 वर्ष। (K=0.09-0.14; tm=10; tmax=23; Fec=150).
Prior r = 0.18, 95% CL = 0.12 - 0.26, Based on 4 full stock assessments.
Fishing Vulnerability (Ref. 59153):  High vulnerability (60 of 100).
Climate Vulnerability (Ref. 125649):  High vulnerability (60 of 100).
Price category (Ref. 80766):   Medium.
Nutrients (Ref. 124155):  Calcium = 9.22 [1.18, 176.74] mg/100g; Iron = 0.628 [0.060, 7.651] mg/100g; Protein = 17.2 [14.4, 19.5] %; Omega3 = 0.571 [0.265, 1.295] g/100g; Selenium = 39.7 [7.5, 183.1] μg/100g; VitaminA = 5.22 [0.40, 59.91] μg/100g; Zinc = 0.47 [0.03, 5.30] mg/100g (wet weight); based on nutrient studies.