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	<dc:identifier>FB-9223</dc:identifier>
	<dc:source>http://www.fishbase.org/summary/SpeciesSummary.php?id=9223</dc:source>
	<dwc:Kingdom>Animalia</dwc:Kingdom>
	<dwc:Phylum>Chordata</dwc:Phylum>
	<dwc:Class>Teleostei</dwc:Class>
	<dwc:Order>Cichliformes</dwc:Order>
	<dwc:Family>Cichlidae</dwc:Family>
	<dwc:Genus>Coptodon</dwc:Genus>
	<dwc:ScientificName>Coptodon margaritacea (Boulenger, 1916)</dwc:ScientificName>
	<commonName xml:lang='fi'>Helmitilapia</commonName>
	<commonName xml:lang=''>珍珠非鯽</commonName>
	<commonName xml:lang=''>珍珠非鲫</commonName>
	<synonym relationship='misapplied name'>Tilapia kottae (non Lönnberg, 1904)</synonym>
	<synonym relationship='synonym'>Tilapia margaritacea Boulenger, 1916</synonym>
	<synonym relationship='misapplied name'>Tilapia tholloni (non Sauvage, 1884)</synonym>
	<dcterms:created>1994-03-25</dcterms:created>
	<dcterms:modified>2024-04-16</dcterms:modified>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=4517&amp;speccode=9223'>Hureau, J.-C.0 La base de données GICIM: Gestion informatisée des collections ichthyologiques du Muséum. p. 225-227. In Atlas Préliminaire des Poissons d'Eau Douce de France. Conseil Supérieur de la Pêche, Ministère de l'Environnement, CEMAGREF et Muséum national d'Histoire naturelle, Paris. (Ref. 4517)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=5163&amp;speccode=9223'>Teugels, G.G. and D.F.E. Thys van den Audenaerde0 Tilapia. p. 482-508. In J. Daget, J.-P. Gosse, G.G. Teugels and D.F.E. Thys van den Audenaerde (eds.) Check-list of the freshwater fishes of Africa (CLOFFA). ISNB, Brussels; MRAC, Tervuren; and ORSTOM, Paris. Vol. 4. (Ref. 5163)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=12818&amp;speccode=9223'>Anonymous0 Fish registrations within the museum database of the Vertebrate Section of the Royal Museum for Central Africa. MRAC, Tervuren, Belgium. (Ref. 12818)</reference>
	<reference isbn='957-8596-59-6' url='http://www.fishbase.org/references/FBRefSummary.php?id=31517&amp;speccode=9223'>Wu, H.L., K.-T. Shao and C.F. Lai (eds.)0 Latin-Chinese dictionary of fishes names. The Sueichan Press, Taiwan. 1028 p. (Ref. 31517)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=31982&amp;speccode=9223'>Anonymous0 Fish collection database of the Natural History Museum, London (formerly British Museum of Natural History (BMNH)). Natural History Museum, London (formerly British Museum of Natural History (BMNH)). (Ref. 31982)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=51086&amp;speccode=9223'>Varjo, M., L. Koli and H. Dahlström0 Kalannimiluettelo (versio 10/03). Suomen Biologian Seura Vanamo Ry. (Ref. 51086)</reference>
	<reference isbn='2.11.086730-2' url='http://www.fishbase.org/references/FBRefSummary.php?id=51193&amp;speccode=9223'>Vivien, J.0 Faune du Cameroun. Guide des mammifères et poissons. GICAM et Ministère de la Coopération et du Développement, Yaoundé/Paris. 271 p. (Ref. 51193)</reference>
	<reference isbn='3-928819-33-X' url='http://www.fishbase.org/references/FBRefSummary.php?id=52307&amp;speccode=9223'>Lamboj, A.0 The cichlid fishes of Western Africa. Birgit Schmettkamp Verlag, Bornheim, Germany. 255 p. (Ref. 52307)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=58108&amp;speccode=9223'>Chinese Academy of Fishery Sciences0 Chinese aquatic germplasm resources database. http://zzzy.fishinfo.cn (Ref. 58108)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=81260&amp;speccode=9223'>Stiassny, M.L.J., A. Lamboj, D. De Weirdt and G.G. Teugels0 Cichlidae. p. 269-403. In M.L.J. Stiassny, G.G Teugels and C.D. Hopkins (eds.) The fresh and brackish water fishes of Lower Guinea, West-Central Africa Volume 2. Coll. faune et flore tropicales 42. Institut de recherche de développement, Paris, France, Muséum national d'histoire naturelle, Paris, France and Musée royal de l'Afrique Central, Tervuren, Belgium, 603p. (Ref. 81260)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=83882&amp;speccode=9223'>FAO-FIES0 Aquatic Sciences and Fisheries Information System (ASFIS) species list. Retrieved from http://www.fao.org/fishery/collection/asfis/en, March 2010. (Ref. 83882)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=90062&amp;speccode=9223'>FAO-FIES0 Aquatic Sciences and Fisheries Information System (ASFIS) species list. Retrieved from http://www.fao.org/fishery/collection/asfis/en, March 2012. (Ref. 90062)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=93285&amp;speccode=9223'>Dunz, A.R. and U.K. Schliewen0 Molecular phylogeny and revised classification of the haplotilapiine cichlid fishes formerly referred to as "&lt;i&gt;Tilapia&lt;/i&gt;". Mol. Phylogenet. Evol. 68(1):64-80. (Ref. 93285)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=95632&amp;speccode=9223'>FAO-FIES0 Aquatic Sciences and Fisheries Information System (ASFIS) species list. Retrieved from http://www.fao.org/fishery/collection/asfis/en, April 2014. (Ref. 95632)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=101110&amp;speccode=9223'>FAO-FIES0 Aquatic Sciences and Fisheries Information System (ASFIS) species list. Retrieved from http://www.fao.org/fishery/collection/asfis/en, [accessed 13/04/2015]. (Ref. 101110)</reference><dataObject>
	<dc:identifier>FB-GeneralDescr-9223</dc:identifier>
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	<agent homepage='' logoURL='' role='recorder'>Pascualita Sa-a</agent>
	<dcterms:created>1994-03-25 00:00:00</dcterms:created>
	<dcterms:modified>2024-04-16 00:00:00</dcterms:modified>
	<dc:title>Biology</dc:title>
	<dc:language></dc:language>
	<license>http://creativecommons.org/licenses/by-nc/3.0/</license>
	<dc:rights></dc:rights>
	<dcterms:rightsHolder>WorldFish Center - FishBase</dcterms:rightsHolder>
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	<audience>General public</audience>
	<dc:source>http://www.fishbase.org/summary/SpeciesSummary.php?id=9223</dc:source>
	<subject>http://rs.tdwg.org/ontology/voc/SPMInfoItems#Distribution</subject>
	<dc:description>Pair-bonding, open (Ref. 52307) substrate brooder (Ref. 52307, 81260), with both parents guarding the brood (Ref. 52307).</dc:description>
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	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=93285&amp;speccode=9223'>Dunz, A.R. and U.K. Schliewen 2013 Molecular phylogeny and revised classification of the haplotilapiine cichlid fishes formerly referred to as "&lt;i&gt;Tilapia&lt;/i&gt;". Mol. Phylogenet. Evol. 68(1):64-80. (Ref. 93285)</reference></dataObject><dataObject>
	<dc:identifier>FB-DiagnosticDescr-9223</dc:identifier>
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	<agent homepage='http://www.africamuseum.be#' logoURL='' role='recorder'>Tobias Musschoot</agent>
	<dcterms:created>2011-03-09 00:00:00</dcterms:created>
	<dcterms:modified>2011-03-09 00:00:00</dcterms:modified>
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	<dc:source>http://www.fishbase.org/summary/SpeciesSummary.php?id=9223</dc:source>
	<dc:description>Diagnosis: 9 or fewer rakers on lower limb of first arch; pearly spots on cheek and opercle; bluish pearly oblique band across dorsal fin from base of first spine to posterior border of soft rays; outer row jaw teeth slender and rather pointed; 2 scale rows on cheek (rarely 3); dorsal fin with 13-15 spines and 10-13 soft rays (total of 25-27 fin rays) (Ref. 81260).Description: head length 33.6-37.9% SL; lower pharyngeal jaw with pointed, recurved anterior teeth and straight tricuspid posterior teeth; ventral keel much shorter than dentigerous plate; 3-6 rows of teeth on oral jaws, outer row teeth biscupid (Ref. 81260).Coloration: body ground color yellow, belly whitish, each scale with a black basal spot; 7-9 vertical bars on upper flanks and caudal peduncle, lending a darker appearance to the dorsum; anteriorly, bars expanded at level of lower lateral line, creating a line of 5-6 black spots at mid-body; dorsum of head, snout and lips blackish; opercle and cheek yellowish; upper lip with bluish hue; many pearly spots or iridophores on the cheek; dorsal fin with fine pearly blue band from base of first spine to extremity of soft dorsal fin; band is only on membrane between rays; fin above blue band is greyish to transparent; the part below is bluish black with small pearly blue spots; caudal fin bluish black with pearly blue lines that split into small lines or spots distally; anal fin bluish black with some pearly blue lines only on the distal part of upper lobe; pelvics with black leading edge, rest of fin grey; pectoral fins transparent (Ref. 81260).</dc:description>
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	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=81260&amp;speccode=9223'>Stiassny, M.L.J., A. Lamboj, D. De Weirdt and G.G. Teugels 2008 Cichlidae. p. 269-403. In M.L.J. Stiassny, G.G Teugels and C.D. Hopkins (eds.) The fresh and brackish water fishes of Lower Guinea, West-Central Africa Volume 2. Coll. faune et flore tropicales 42. Institut de recherche de développement, Paris, France, Muséum national d'histoire naturelle, Paris, France and Musée royal de l'Afrique Central, Tervuren, Belgium, 603p. (Ref. 81260)</reference></dataObject><dataObject>
	<dc:identifier>FB-Morphology-9223</dc:identifier>
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	<agent homepage='http://www.africamuseum.be#' logoURL='' role='recorder'>Tobias Musschoot</agent>
	<dcterms:created>2011-03-09 00:00:00</dcterms:created>
	<dcterms:modified>2011-03-09 00:00:00</dcterms:modified>
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	<dcterms:rightsHolder>WorldFish Center - FishBase</dcterms:rightsHolder>
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	<audience>General public</audience>
	<dc:source>http://www.fishbase.org/summary/SpeciesSummary.php?id=9223</dc:source>
	<subject>http://rs.tdwg.org/ontology/voc/SPMInfoItems#Size</subject>
	<dc:description>Dorsal spines (total): 13 - 15; Dorsal soft rays (total): 10 - 13;  Analspines: 3; Analsoft rays: 8 - 10</dc:description>
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	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=81260&amp;speccode=9223'>Stiassny, M.L.J., A. Lamboj, D. De Weirdt and G.G. Teugels 2008 Cichlidae. p. 269-403. In M.L.J. Stiassny, G.G Teugels and C.D. Hopkins (eds.) The fresh and brackish water fishes of Lower Guinea, West-Central Africa Volume 2. Coll. faune et flore tropicales 42. Institut de recherche de développement, Paris, France, Muséum national d'histoire naturelle, Paris, France and Musée royal de l'Afrique Central, Tervuren, Belgium, 603p. (Ref. 81260)</reference></dataObject><dataObject>
	<dc:identifier>FB-Distribution-9223</dc:identifier>
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	<agent homepage='' logoURL='' role='recorder'>Pascualita Sa-a</agent>
	<dcterms:created>1994-03-25 00:00:00</dcterms:created>
	<dcterms:modified>2025-08-12 00:00:00</dcterms:modified>
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	<dc:language></dc:language>
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	<dcterms:bibliographicCitation></dcterms:bibliographicCitation>
	<audience>General public</audience>
	<dc:source>http://www.fishbase.org/summary/SpeciesSummary.php?id=9223</dc:source>
	<subject>http://rs.tdwg.org/ontology/voc/SPMInfoItems#Threats</subject>
	<dc:description>Africa:  known only from the Nyong basin in Cameroon (Ref. 52307, 81260).</dc:description>
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	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=5163&amp;speccode=9223'>Teugels, G.G. and D.F.E. Thys van den Audenaerde 1991 Tilapia. p. 482-508. In J. Daget, J.-P. Gosse, G.G. Teugels and D.F.E. Thys van den Audenaerde (eds.) Check-list of the freshwater fishes of Africa (CLOFFA). ISNB, Brussels; MRAC, Tervuren; and ORSTOM, Paris. Vol. 4. (Ref. 5163)</reference></dataObject><dataObject>
	<dc:identifier>FB-Size-9223</dc:identifier>
	<dataType>http://purl.org/dc/dcmitype/Text</dataType>
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	<agent homepage='' logoURL='' role='recorder'>Pascualita Sa-a</agent>
	<dcterms:created>1994-03-25 00:00:00</dcterms:created>
	<dcterms:modified>2024-04-16 00:00:00</dcterms:modified>
	<dc:title>Max. size</dc:title>
	<dc:language></dc:language>
	<license>http://creativecommons.org/licenses/by-nc/3.0/</license>
	<dc:rights></dc:rights>
	<dcterms:rightsHolder>WorldFish Center - FishBase</dcterms:rightsHolder>
	<dcterms:bibliographicCitation></dcterms:bibliographicCitation>
	<audience>General public</audience>
	<dc:source>http://www.fishbase.org/summary/SpeciesSummary.php?id=9223</dc:source>
	<subject>http://rs.tdwg.org/ontology/voc/SPMInfoItems#Habitat</subject>
	<dc:description>17.5 cm TL (male/unsexed; (Ref. 5163))</dc:description>
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	<thumbnailURL></thumbnailURL>
	<location></location>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=5163&amp;speccode=9223'>Teugels, G.G. and D.F.E. Thys van den Audenaerde 1991 Tilapia. p. 482-508. In J. Daget, J.-P. Gosse, G.G. Teugels and D.F.E. Thys van den Audenaerde (eds.) Check-list of the freshwater fishes of Africa (CLOFFA). ISNB, Brussels; MRAC, Tervuren; and ORSTOM, Paris. Vol. 4. (Ref. 5163)</reference></dataObject><dataObject>
	<dc:identifier>FB-Threats-2</dc:identifier>
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	<agent homepage='' logoURL='' role='recorder'>Belen Acosta</agent>
	<dcterms:created>1990-10-18 00:00:00</dcterms:created>
	<dcterms:modified>2024-03-07 00:00:00</dcterms:modified>
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	<audience>General public</audience>
	<dc:source>http://www.fishbase.org/summary/SpeciesSummary.php?id=2</dc:source>
	<subject>http://rs.tdwg.org/ontology/voc/SPMInfoItems#DiagnosticDescription</subject>
	<dc:description>Least Concern (LC) , IUCN Grouper and Wrasse Specialist Group</dc:description>
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	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=57073'>IUCN 2006 2006 IUCN red list of threatened species. www.iucnredlist.org. Downloaded July 2006.</reference></dataObject><dataObject>
	<dc:identifier>FB-Habitat-9223</dc:identifier>
	<dataType>http://purl.org/dc/dcmitype/Text</dataType>
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	<agent homepage='' logoURL='' role='recorder'>Pascualita Sa-a</agent>
	<dcterms:created>1994-03-25 00:00:00</dcterms:created>
	<dcterms:modified>2024-04-16 00:00:00</dcterms:modified>
	<dc:title>Environment</dc:title>
	<dc:language></dc:language>
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	<dc:source>http://www.fishbase.org/summary/SpeciesSummary.php?id=9223</dc:source>
	<dc:description>benthopelagic; freshwater</dc:description>
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