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	<dc:identifier>FB-8968</dc:identifier>
	<dc:source>http://www.fishbase.org/summary/SpeciesSummary.php?id=8968</dc:source>
	<dwc:Kingdom>Animalia</dwc:Kingdom>
	<dwc:Phylum>Chordata</dwc:Phylum>
	<dwc:Class>Elasmobranchii</dwc:Class>
	<dwc:Order>Myliobatiformes</dwc:Order>
	<dwc:Family>Dasyatidae</dwc:Family>
	<dwc:Genus>Fontitrygon</dwc:Genus>
	<dwc:ScientificName>Fontitrygon ukpam (Smith, 1863)</dwc:ScientificName>
	<commonName xml:lang='en'>Pincushion ray</commonName>
	<commonName xml:lang='en'>Thorny freshwater stingray</commonName>
	<commonName xml:lang='et'>Ukpam-astelrai</commonName>
	<commonName xml:lang=''>Speldenkussenpijlstaartrog</commonName>
	<commonName xml:lang=''>尤克魟</commonName>
	<synonym relationship='misapplied name'>Dasyatis centroura (non Mitchill, 1815)</synonym>
	<synonym relationship='synonym'>Dasyatis ukpam (Smith, 1863)</synonym>
	<synonym relationship='synonym'>Hemitrygon ukpam Smith, 1863</synonym>
	<synonym relationship='synonym'>Trygon ukpam (Smith, 1863)</synonym>
	<synonym relationship='synonym'>Urogymnus ukpam (Smith, 1863)</synonym>
	<dcterms:created>1994-03-05</dcterms:created>
	<dcterms:modified>2024-07-18</dcterms:modified>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=205&amp;speccode=8968'>Breder, C.M. and D.E. Rosen0 Modes of reproduction in fishes. T.F.H. Publications, Neptune City, New Jersey. 941 p. (Ref. 205)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=3497&amp;speccode=8968'>Compagno, L.J.V. and T.R. Roberts0 Dasyatidae. p. 4-5. In J. Daget, J.P. Gosse and D.F.E. Thys van den Audenaerde (eds.) Check-list of the freshwater fishes of Africa (CLOFFA). ORSTOM, Paris and MRAC, Tervuren. Vol. 1. (Ref. 3497)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=4438&amp;speccode=8968'>Capapé, C. and M. Desoutter0 Dasyatidae. p. 59-63. In J.C. Quero, J.C. Hureau, C. Karrer, A. Post and L. Saldanha (eds.) Check-list of the fishes of the eastern tropical Atlantic (CLOFETA). JNICT, Lisbon; SEI, Paris; and UNESCO, Paris. Vol. 1. (Ref. 4438)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=4517&amp;speccode=8968'>Hureau, J.-C.0 La base de données GICIM: Gestion informatisée des collections ichthyologiques du Muséum. p. 225-227. In Atlas Préliminaire des Poissons d'Eau Douce de France. Conseil Supérieur de la Pêche, Ministère de l'Environnement, CEMAGREF et Muséum national d'Histoire naturelle, Paris. (Ref. 4517)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=7397&amp;speccode=8968'>Séret, B.0 Dasyatidae. p. 62-75. In C. Levêque, D. Paugy, and G.G. Teugels (eds.) Faune des poissons d'eaux douces et saumâtres d'Afrique de l'Ouest Tome 1. Coll. Faune Tropicale n° 28. Musée Royal de l'Afrique Centrale, Tervuren, Belgique and O.R.S.T.O.M., Paris, France, 384 p. (Ref. 7397)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=12818&amp;speccode=8968'>Anonymous0 Fish registrations within the museum database of the Vertebrate Section of the Royal Museum for Central Africa. MRAC, Tervuren, Belgium. (Ref. 12818)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=26277&amp;speccode=8968'>Compagno, L.J.V. and T.R. Roberts0 Marine and freshwater stingrays (Dasyatidae) of West Africa, with description of a new species. Proc. Calif. Acad. Sci. 43(18):283-300. (Ref. 26277)</reference>
	<reference isbn='957-8596-59-6' url='http://www.fishbase.org/references/FBRefSummary.php?id=31517&amp;speccode=8968'>Wu, H.L., K.-T. Shao and C.F. Lai (eds.)0 Latin-Chinese dictionary of fishes names. The Sueichan Press, Taiwan. 1028 p. (Ref. 31517)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=31982&amp;speccode=8968'>Anonymous0 Fish collection database of the Natural History Museum, London (formerly British Museum of Natural History (BMNH)). Natural History Museum, London (formerly British Museum of Natural History (BMNH)). (Ref. 31982)</reference>
	<reference isbn='0-8018-6048-2' url='http://www.fishbase.org/references/FBRefSummary.php?id=35766&amp;speccode=8968'>Compagno, L.J.V.0 Checklist of living elasmobranchs. p. 471-498. In W.C. Hamlett (ed.) Sharks, skates, and rays: the biology of elasmobranch fishes. Johns Hopkins University Press, Maryland. (Ref. 35766)</reference>
	<reference isbn='2-8317-0564-9' url='http://www.fishbase.org/references/FBRefSummary.php?id=36508&amp;speccode=8968'>Hilton-Taylor, C.0 2000 IUCN red list of threatened species. IUCN, Gland, Switzerland and Cambridge, UK. xviii + 61 p. (with 1 CD-ROM). (Ref. 36508)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=38732&amp;speccode=8968'>Anonymous0 Fish collection database of the National Museum of Natural History (Smithsonian Institution). Smithsonian Institution-Division of Fishes. (Ref. 38732)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=38974&amp;speccode=8968'>Taniuchi, T.0 Occurrence of two species of stingrays of the genus &lt;i&gt;Dasyatis&lt;/i&gt; (Chondrichthyes) in the Sanaga basin, Cameroun. Environ. Biol. Fishes 31(1):95-100. (Ref. 38974)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=43454&amp;speccode=8968'>Bor, P.H.F.0 Nederlandse naamlijst van de recente haaien en roggen (Chondrichthyes: Elasmobranchii) van de wereld. World Wide Web electronic publication www.rajidae.tmfweb.nl, version (05/2002). (Ref. 43454)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=50449&amp;speccode=8968'>Dulvy, N.K. and J.D. Reynolds0 Evolutionary transitions among egg-laying, live-bearing and maternal inputs in sharks and rays. Proc. R. Soc. Lond., Ser. B: Biol. Sci. 264:1309-1315. (Ref. 50449)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=58108&amp;speccode=8968'>Chinese Academy of Fishery Sciences0 Chinese aquatic germplasm resources database. http://zzzy.fishinfo.cn (Ref. 58108)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=78745&amp;speccode=8968'>Albaret, J.-J., M. Simier, F.S. Darboe, J.-M. Ecoutin, J. Raffray and L.T. de Morais0 Fish diversity and distribution in the Gambia estuary, West Africa, in relation to environmental variables. Aquat. Living Resour. 17:35-46. (Ref. 78745)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=81259&amp;speccode=8968'>Séret, B.0 Dasyatidae. p. 81-96. In D. Paugy, C. Lévêque and G.G Teugels (eds.) The fresh and brackish water fishes of West Africa Volume 1. Collection Faune et Flore Tropicales 40. Institut de recherche de développement, Paris, France, Muséum national d'histoire naturelle, Paris, France and Musée royal de l'Afrique Central, Tervuren, Belgium, 457p. (Ref. 81259)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=81625&amp;speccode=8968'>de Carvalho, M.R., B. Séret and J.D. McEachran0 Dasyatidae. p. 154-164. In M.L.J. Stiassny, G.G. Teugels and C.D. Hopkins (eds.) The fresh and brackish water fishes of Lower Guinea, West-Central Africa. Volume I. Collection Faune et Flore tropicales 42. Institut de Recherche pour le Développement, Paris, France, Muséum National d’Histoire Naturelle, Paris, France, and Musée Royal de l’Afrique Centrale, Tervuren, Belgium. 800 pp. (Ref. 81625)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=83882&amp;speccode=8968'>FAO-FIES0 Aquatic Sciences and Fisheries Information System (ASFIS) species list. Retrieved from http://www.fao.org/fishery/collection/asfis/en, March 2010. (Ref. 83882)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=90062&amp;speccode=8968'>FAO-FIES0 Aquatic Sciences and Fisheries Information System (ASFIS) species list. Retrieved from http://www.fao.org/fishery/collection/asfis/en, March 2012. (Ref. 90062)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=95632&amp;speccode=8968'>FAO-FIES0 Aquatic Sciences and Fisheries Information System (ASFIS) species list. Retrieved from http://www.fao.org/fishery/collection/asfis/en, April 2014. (Ref. 95632)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=97748&amp;speccode=8968'>Smith, J.A.0 Notice of the ukpam, a large species (probably new) of sting ray (&lt;i&gt;Trygon&lt;/i&gt;, Cuv.), found in the Old Calabar River, Africa. Proc. R. Phys. Soc. Edinb. 2:64-69. (Ref. 97748)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=101110&amp;speccode=8968'>FAO-FIES0 Aquatic Sciences and Fisheries Information System (ASFIS) species list. Retrieved from http://www.fao.org/fishery/collection/asfis/en, [accessed 13/04/2015]. (Ref. 101110)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=105471&amp;speccode=8968'>eElurikkus0 eElurikkus koondab ühte portaali Eesti eluslooduse andmed. http://elurikkus.ut.ee/ (Ref. 105471)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=114953&amp;speccode=8968'>Last, P.R., W.T. White, M.R. de Carvalho, B. Séret, M.F.W. Stehmann and G.J.P. Naylor0 Rays of the world. CSIRO Publishing, Comstock Publishing Associates. i-ix + 1-790. (Ref. 114953)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=115257&amp;speccode=8968'>FAO-FIES0 Aquatic Sciences and Fisheries Information System (ASFIS) species list. Retrieved from http://www.fao.org/fishery/collection/asfis/en (accessed 08/06/2017). (Ref. 115257)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=118272&amp;speccode=8968'>FAO-FIES0 Aquatic Sciences and Fisheries Information System (ASFIS) species list. Retrieved from htttps://www.fao.org/fishery/collection/asfis/en [accessed May, 2018] (Ref. 118272)</reference><dataObject>
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	<dcterms:created>1994-03-05 00:00:00</dcterms:created>
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	<dc:title>Biology</dc:title>
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	<audience>General public</audience>
	<dc:source>http://www.fishbase.org/summary/SpeciesSummary.php?id=8968</dc:source>
	<subject>http://rs.tdwg.org/ontology/voc/SPMInfoItems#TrophicStrategy</subject>
	<dc:description>Benthic species mostly in lakes and rives, found more than 240 km upstream from the sea (possibly also coastal).  The life history and present status of populations in West Africa is unknown.  Type specimens were born after capture of the mother; few specimens exist (Ref. 114953).  Ovoviviparous (Ref. 50449).</dc:description>
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	<reference isbn='0-8018-6048-2' url='http://www.fishbase.org/references/FBRefSummary.php?id=35766&amp;speccode=8968'>Compagno, L.J.V. 1999 Checklist of living elasmobranchs. p. 471-498. In W.C. Hamlett (ed.) Sharks, skates, and rays: the biology of elasmobranch fishes. Johns Hopkins University Press, Maryland. (Ref. 35766)</reference></dataObject><dataObject>
	<dc:identifier>FB-DiagnosticDescr-8968</dc:identifier>
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	<agent homepage='http://www.africamuseum.be#' logoURL='' role='recorder'>Gert Boden</agent>
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	<subject>http://rs.tdwg.org/ontology/voc/SPMInfoItems#Size</subject>
	<dc:description>Diagnosis: Dasyatis ukpam is a very large and thick-bodied freshwater dasyatid (Ref. 26277). Entire dorsal surface of disc covered with stout-spined denticles, smooth in newborn (Ref. 26277). Sting greatly reduced in size or absent (Ref. 26277).Description: A very large stingray with a thick and almost circular disc, with broadly rounded corners and about as wide as long, and a scarcely distinct snout (Ref. 81259, 81625). Disc oval, very thick; medial lobe narrow-based and short, its anterior margin broadly convex; disc depth 13.3-15.7% of disc width; dorsal surface of disc very rough, entirely covered with rounded or heart-shaped stout-spined dermal denticles, which are particularly concentrated on the centre of the disc (Ref. 26277, 81259). One or two pearly tubercles sometimes present, but especially an irregular mid-dorsal row of large, conical pointed thorny tubercles and a few similar ones scattered over the pectoralfins (Ref. 81259). Mouth slightly arched, with 38-46/38-48 rows of teeth set in a pavement pattern, and 4 or 5 elongated finger-like mouth papillae (Ref. 7397, 81259, 81625). Nasal curtain fringed along its posterior margin, eyes relatively small and slightly elevated, spiracles large (Ref. 81625). Pelvic fins almost entirely covered by disc dorsally (Ref. 81625). Total pectoral radials 142-149 (Ref. 26277). Tail long and very spiny, rather stout and circular in cross-section at base; sting reduced or absent, dorsal ridge absent and ventral fold reduced (Ref. 81259, 81625).Colouration: Disc and pelvic fins dark brown or gray-brown above, without spots or prominent markings; belly white except for broad dark margins; tail blackish except for underside of its base (Ref. 26277, 81259, 81625).</dc:description>
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	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=26277&amp;speccode=8968'>Compagno, L.J.V. and T.R. Roberts 1984 Marine and freshwater stingrays (Dasyatidae) of West Africa, with description of a new species. Proc. Calif. Acad. Sci. 43(18):283-300. (Ref. 26277)</reference></dataObject><dataObject>
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	<agent homepage='http://www.africamuseum.be#' logoURL='' role='recorder'>Gert Boden</agent>
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	<subject>http://rs.tdwg.org/ontology/voc/SPMInfoItems#Threats</subject>
	<dc:description>Vertebrae: 108 - 122</dc:description>
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	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=26277&amp;speccode=8968'>Compagno, L.J.V. and T.R. Roberts 1984 Marine and freshwater stingrays (Dasyatidae) of West Africa, with description of a new species. Proc. Calif. Acad. Sci. 43(18):283-300. (Ref. 26277)</reference></dataObject><dataObject>
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	<subject>http://rs.tdwg.org/ontology/voc/SPMInfoItems#Habitat</subject>
	<dc:description>Africa:  rivers and lakes in western Africa (Ref. 81259, 81625).</dc:description>
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	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=81259&amp;speccode=8968'>Séret, B. 2003 Dasyatidae. p. 81-96. In D. Paugy, C. Lévêque and G.G Teugels (eds.) The fresh and brackish water fishes of West Africa Volume 1. Collection Faune et Flore Tropicales 40. Institut de recherche de développement, Paris, France, Muséum national d'histoire naturelle, Paris, France and Musée royal de l'Afrique Central, Tervuren, Belgium, 457p. (Ref. 81259)</reference></dataObject><dataObject>
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	<agent homepage='' logoURL='' role='recorder'>Susan M. Luna</agent>
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	<dc:source>http://www.fishbase.org//Reproduction/FishReproSummary.php?ID=8968&amp;StockCode=9289</dc:source>
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	<dc:description>Exhibit ovoviparity (aplacental viviparity), with embryos feeding initially on yolk, then receiving additional nourishment from the mother by indirect absorption of uterine fluid enriched with mucus, fat or protein through specialised structures (Ref. 50449). Distinct pairing with embrace (Ref. 205).</dc:description>
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	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=205&amp;speccode=8968'>Breder, C.M. and D.E. Rosen 1966 Modes of reproduction in fishes. T.F.H. Publications, Neptune City, New Jersey. 941 p. (Ref. 205)</reference></dataObject><dataObject>
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	<agent homepage='' logoURL='' role='recorder'>Crispina B. Binohlan</agent>
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	<dc:title>Max. size</dc:title>
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	<subject>http://rs.tdwg.org/ontology/voc/SPMInfoItems#Distribution</subject>
	<dc:description>120 cm WD (male/unsexed; (Ref. 81259))</dc:description>
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	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=81259&amp;speccode=8968'>Séret, B. 2003 Dasyatidae. p. 81-96. In D. Paugy, C. Lévêque and G.G Teugels (eds.) The fresh and brackish water fishes of West Africa Volume 1. Collection Faune et Flore Tropicales 40. Institut de recherche de développement, Paris, France, Muséum national d'histoire naturelle, Paris, France and Musée royal de l'Afrique Central, Tervuren, Belgium, 457p. (Ref. 81259)</reference></dataObject><dataObject>
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	<agent homepage='' logoURL='' role='recorder'>Belen Acosta</agent>
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	<dcterms:modified>2024-03-07 00:00:00</dcterms:modified>
	<dc:title></dc:title>
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	<dc:rights></dc:rights>
	<dcterms:bibliographicCitation></dcterms:bibliographicCitation>
	<dc:source>http://www.fishbase.org/summary/SpeciesSummary.php?id=2</dc:source>
	<dc:description>Least Concern (LC) , IUCN Grouper and Wrasse Specialist Group</dc:description>
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	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=57073'>IUCN 2006 2006 IUCN red list of threatened species. www.iucnredlist.org. Downloaded July 2006.</reference></dataObject><dataObject>
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	<agent homepage='' logoURL='' role='recorder'>Crispina B. Binohlan</agent>
	<dcterms:created>1994-03-05 00:00:00</dcterms:created>
	<dcterms:modified>2024-07-18 00:00:00</dcterms:modified>
	<dc:title>Environment</dc:title>
	<dc:language></dc:language>
	<license>http://creativecommons.org/licenses/by-nc/3.0/</license>
	<dc:rights></dc:rights>
	<dcterms:rightsHolder>WorldFish Center - FishBase</dcterms:rightsHolder>
	<dcterms:bibliographicCitation></dcterms:bibliographicCitation>
	<audience>General public</audience>
	<dc:source>http://www.fishbase.org/summary/SpeciesSummary.php?id=8968</dc:source>
	<subject>http://rs.tdwg.org/ontology/voc/SPMInfoItems#Size</subject>
	<dc:description>demersal; freshwater; brackish</dc:description>
	<mediaURL></mediaURL>
	<thumbnailURL></thumbnailURL>
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