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	<dc:identifier>FB-66292</dc:identifier>
	<dc:source>http://www.fishbase.org/summary/SpeciesSummary.php?id=66292</dc:source>
	<dwc:Kingdom>Animalia</dwc:Kingdom>
	<dwc:Phylum>Chordata</dwc:Phylum>
	<dwc:Class>Teleostei</dwc:Class>
	<dwc:Order>Callionymiformes</dwc:Order>
	<dwc:Family>Callionymidae</dwc:Family>
	<dwc:Genus>Tonlesapia</dwc:Genus>
	<dwc:ScientificName>Tonlesapia amnica Ng &amp; Rainboth, 2011</dwc:ScientificName>
	<commonName xml:lang='en'>Sculpted lanternshark</commonName>
	<commonName xml:lang='et'>Raidhammas-tumehai</commonName>
	<synonym relationship='misapplied name'>Etmopterus brachyurus (non Smith &amp; Radcliffe, 1912)</synonym>
	<synonym relationship='misapplied name'>Etmopterus lucifer (non Jordan &amp; Snyder, 1902)</synonym>
	<synonym relationship='ambiguous synonym'>Etmopterus sp. Not applicable</synonym>
	<synonym relationship='misapplied name'>Spinax lucifer (non Jordan &amp; Snyder, 1902)</synonym>
	<dcterms:created>2011-10-07</dcterms:created>
	<dcterms:modified>2024-03-18</dcterms:modified>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=205&amp;speccode=66301'>Breder, C.M. and D.E. Rosen0 Modes of reproduction in fishes. T.F.H. Publications, Neptune City, New Jersey. 941 p. (Ref. 205)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=1953&amp;speccode=66301'>Smith, J.L.B.0 The sea fishes of southern Africa. Central News Agency Ltd., Cape Town. 550 p. (Ref. 1953)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=5578&amp;speccode=66301'>Compagno, L.J.V., D.A. Ebert and M.J. Smale0 Guide to the sharks and rays of southern Africa. New Holland (Publ.) Ltd., London. 158 p. (Ref. 5578)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=6577&amp;speccode=66301'>Bass, A.J., L.J.V. Compagno and P.C. Heemstra0 Squalidae. p. 49-62. In M.M. Smith and P.C. Heemstra (eds.) Smiths' sea fishes. Springer-Verlag, Berlin. (Ref. 6577)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=12473&amp;speccode=66301'>Ebert, D.A., L.J.V. Compagno and P.D. Cowley0 A preliminary investigation of the feeding ecology of squaloid sharks off the west coast of southern Africa. S. Afr. J. mar Sci. 12:601-609. (Ref. 12473)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=27644&amp;speccode=66301'>Compagno, L.V.J., D.E. Ebert and P.D. Cowley0 Distribution of offshore demersal cartilaginous fish (Class Chondrichthyes) off the west coast of southern Africa, with notes on their systematics. S. Afr. J. Mar. Sci. 11:43-139. (Ref. 27644)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=32002&amp;speccode=66301'>Norman, J.R.0 Coast fishes. Part I. The South Atlantic. p. 57-58. In Discovery Report, vol. XII. (Ref. 32002)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=36031&amp;speccode=66301'>Barnard, K.H.0 A monograph of the marine fishes of South Africa. Part I. Ann. S. Afr. Mus. 21:1-418. (Ref. 36031)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=48413&amp;speccode=66301'>Bigelow, H.B. and W.C. Schroeder0 A study of the sharks of the suborder Squaloidea. Bull. Mus. Comp. Zool. 117(1):1-150. (Ref. 48413)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=58085&amp;speccode=66301'>Compagno, L.J.V., D. Dando and S. Fowler0 Sharks of the world. Princeton field guides. Harper Collins Publishing Ltd., London. 368 p. (Ref. 58085)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=87957&amp;speccode=66301'>Ebert, D.A., Compagno, L.J.V. and M.J. De Vries0 A New Lanternshark (Squaliformes: Etmopteridae: &lt;i&gt;Etmopterus&lt;/i&gt;) from Southern Africa. Copeia 2011(3):379-384. (Ref. 87957)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=88004&amp;speccode=66301'>Compagno, L.J.V.0 An overview of chondrichthyan systematics and biodiversity in southern Africa. Transactions of the Royal Society of South Africa 54:75-120. (Ref. 88004)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=105471&amp;speccode=66301'>eElurikkus0 eElurikkus koondab ühte portaali Eesti eluslooduse andmed. http://elurikkus.ut.ee/ (Ref. 105471)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=106604&amp;speccode=66301'>Weigmann, S.0 Annotated checklist of the living sharks, batoids and chimaeras (Chondrichthyes) of the world, with a focus on biogeographical diversity. J. Fish Biol. 88(1):1-201. (Ref. 106604)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=115257&amp;speccode=66301'>FAO-FIES0 Aquatic Sciences and Fisheries Information System (ASFIS) species list. Retrieved from http://www.fao.org/fishery/collection/asfis/en (accessed 08/06/2017). (Ref. 115257)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=119149&amp;speccode=66301'>Ebert, D.A. and K.E. Van Heels0 &lt;i&gt;Etmopterus marshae&lt;/i&gt; sp. nov, a new lanternshark (Squaliformes: Etmopteridae) from the Philippine Islands, with a revised key to the &lt;i&gt;Etmopterus lucifer&lt;/i&gt; clade. Zootaxa 4508(2):197-210. (Ref. 119149)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=124468&amp;speccode=66301'>Ebert, D.A., R.W. Leslie and S. Weigmann0 &lt;i&gt;Etmopterus brosei&lt;/i&gt; sp. nov.: a new lanternshark (Squaliformes: Etmopteridae) from the southeastern Atlantic and southwestern Indian oceans, with a revised key to the &lt;i&gt;Etmopterus lucifer&lt;/i&gt; clade. Marine Biodiversity 51(53):1-17. (Ref. 124468)</reference><dataObject>
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	<agent homepage='' logoURL='' role='recorder'>Frédéric Busson</agent>
	<dcterms:created>2011-10-14 23:02:11</dcterms:created>
	<dcterms:modified>2024-03-18 00:00:00</dcterms:modified>
	<dc:title>Biology</dc:title>
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	<dc:source>http://www.fishbase.org/summary/SpeciesSummary.php?id=66301</dc:source>
	<dc:description>This species is fairly common along the upper continental slopes between Namibia and southern Mozambique (Ref. 87957); upper slope, on or near the bottom (Ref. 5578).  Observed mostly at depths between 450 and 900 m (Ref. 27644), but with records as shallow as 240 m (in Bass et al., 1976: listed as E. lucifer) (Ref. 87957).  Feeds mainly on mid-water bony fish, also squid, octopus, shrimp and brittle stars (Ref. 5578).</dc:description>
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	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=87957&amp;speccode=66301'>Ebert, D.A., Compagno, L.J.V. and M.J. De Vries 2011 A New Lanternshark (Squaliformes: Etmopteridae: &lt;i&gt;Etmopterus&lt;/i&gt;) from Southern Africa. Copeia 2011(3):379-384. (Ref. 87957)</reference></dataObject><dataObject>
	<dc:identifier>FB-DiagnosticDescr-66301</dc:identifier>
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	<agent homepage='' logoURL='' role='recorder'>Frédéric Busson</agent>
	<dcterms:created>2011-10-15 00:00:00</dcterms:created>
	<dcterms:modified>2017-11-15 00:00:00</dcterms:modified>
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	<dc:source>http://www.fishbase.org/summary/SpeciesSummary.php?id=66301</dc:source>
	<dc:description>Etmopterus sculptus is a moderately large, stout, species of linear-denticled Etmopterus, which separates it from most other members of this genus, and can be distinguished from its three closest congeners within the E. Lucifer-group, E. brachyurus, E. lucifer, and E. molleri, by the length of its anterior flank markings being slightly longer than its posterior branch. The posterior branch of the flank markings on E. brachyurus and E. molleri are shorter relative to the anterior branch, and in E. lucifer, the anterior branch is nearly twice the length of the posterior branch. The new species can also be distinguished from its closest congeners by the arrangement of dermal denticles that extend over most of the upper and lower body surface. Etmopterus sculptus possesses non-overlapping denticles, giving it a rougher, more sculpted texture than these other aforementioned species. The arrangement of the dermal denticles onthe ventral head surface is also a good character for separating these species, with E. sculptus having uniformly distributed denticles while the ventral surface of E. Lucifer and E. molleri is sparsely covered with denticles. The ventral surface of the head of E. brachyurus typically lacks dermal denticles (Ref. 87957).</dc:description>
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	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=87957&amp;speccode=66301'>Ebert, D.A., Compagno, L.J.V. and M.J. De Vries 2011 A New Lanternshark (Squaliformes: Etmopteridae: &lt;i&gt;Etmopterus&lt;/i&gt;) from Southern Africa. Copeia 2011(3):379-384. (Ref. 87957)</reference></dataObject><dataObject>
	<dc:identifier>FB-Distribution-66301</dc:identifier>
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	<agent homepage='' logoURL='' role='recorder'>Frédéric Busson</agent>
	<dcterms:created>2011-10-14 23:29:47</dcterms:created>
	<dcterms:modified>2025-11-05 00:00:00</dcterms:modified>
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	<dc:source>http://www.fishbase.org/summary/SpeciesSummary.php?id=66301</dc:source>
	<dc:description>Southeast Atlantic and Southwest Indian Ocean: Namibia to southern Mozambique,and Madagascar Ridge.</dc:description>
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	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=124468&amp;speccode=66301'>Ebert, D.A., R.W. Leslie and S. Weigmann 2021 &lt;i&gt;Etmopterus brosei&lt;/i&gt; sp. nov.: a new lanternshark (Squaliformes: Etmopteridae) from the southeastern Atlantic and southwestern Indian oceans, with a revised key to the &lt;i&gt;Etmopterus lucifer&lt;/i&gt; clade. Marine Biodiversity 51(53):1-17. (Ref. 124468)</reference></dataObject><dataObject>
	<dc:identifier>FB-Uses-66301</dc:identifier>
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	<agent homepage='' logoURL='' role='recorder'>Frédéric Busson</agent>
	<dcterms:created>2011-10-14 23:02:11</dcterms:created>
	<dcterms:modified>2024-03-18 00:00:00</dcterms:modified>
	<dc:title>Importance</dc:title>
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	<dc:source>http://www.fishbase.org/summary/SpeciesSummary.php?id=66301</dc:source>
	<dc:description>fisheries: of no interest</dc:description>
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	<dc:identifier>FB-LifeCycle-66301</dc:identifier>
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	<agent homepage='' logoURL='' role='recorder'>Cristina V. Garilao</agent>
	<dcterms:created>2011-10-14 23:02:11</dcterms:created>
	<dcterms:modified>2024-03-18 00:00:00</dcterms:modified>
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	<dc:source>http://www.fishbase.org//Reproduction/FishReproSummary.php?ID=66301&amp;StockCode=56961</dc:source>
	<dc:description>Bears 2 young (Ref. 5578). Distinct pairing with embrace (Ref. 205).</dc:description>
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	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=5578&amp;speccode=66301'>Compagno, L.J.V., D.A. Ebert and M.J. Smale 1989 Guide to the sharks and rays of southern Africa. New Holland (Publ.) Ltd., London. 158 p. (Ref. 5578)</reference></dataObject><dataObject>
	<dc:identifier>FB-Size-66301</dc:identifier>
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	<agent homepage='' logoURL='' role='recorder'>Frédéric Busson</agent>
	<dcterms:created>2011-10-14 23:02:11</dcterms:created>
	<dcterms:modified>2024-03-18 00:00:00</dcterms:modified>
	<dc:title>Max. size</dc:title>
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	<dc:source>http://www.fishbase.org/summary/SpeciesSummary.php?id=66301</dc:source>
	<dc:description>59.0 cm TL (male/unsexed; (Ref. 106604)); 51.5 cm TL (female)</dc:description>
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	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=106604&amp;speccode=66301'>Weigmann, S. 2016 Annotated checklist of the living sharks, batoids and chimaeras (Chondrichthyes) of the world, with a focus on biogeographical diversity. J. Fish Biol. 88(1):1-201. (Ref. 106604)</reference></dataObject><dataObject>
	<dc:identifier>FB-Threats-2</dc:identifier>
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	<agent homepage='' logoURL='' role='recorder'>Belen Acosta</agent>
	<dcterms:created>1990-10-18 00:00:00</dcterms:created>
	<dcterms:modified>2024-03-07 00:00:00</dcterms:modified>
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	<dc:source>http://www.fishbase.org/summary/SpeciesSummary.php?id=2</dc:source>
	<dc:description>Least Concern (LC) , IUCN Grouper and Wrasse Specialist Group</dc:description>
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	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=57073'>IUCN 2006 2006 IUCN red list of threatened species. www.iucnredlist.org. Downloaded July 2006.</reference></dataObject><dataObject>
	<dc:identifier>FB-Habitat-66301</dc:identifier>
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	<agent homepage='' logoURL='' role='recorder'>Frédéric Busson</agent>
	<dcterms:created>2011-10-14 23:02:11</dcterms:created>
	<dcterms:modified>2024-03-18 00:00:00</dcterms:modified>
	<dc:title>Environment</dc:title>
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	<dc:source>http://www.fishbase.org/summary/SpeciesSummary.php?id=66301</dc:source>
	<dc:description>bathydemersal; marine; depth range 240 - 1023 m (Ref. 106604)</dc:description>
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	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=106604&amp;speccode=66301'>Weigmann, S. 2016 Annotated checklist of the living sharks, batoids and chimaeras (Chondrichthyes) of the world, with a focus on biogeographical diversity. J. Fish Biol. 88(1):1-201. (Ref. 106604)</reference></dataObject>
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