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	<dc:identifier>FB-604</dc:identifier>
	<dc:source>http://www.fishbase.org/summary/SpeciesSummary.php?id=604</dc:source>
	<dwc:Kingdom>Animalia</dwc:Kingdom>
	<dwc:Phylum>Chordata</dwc:Phylum>
	<dwc:Class>Teleostei</dwc:Class>
	<dwc:Order>Clupeiformes</dwc:Order>
	<dwc:Family>Engraulidae</dwc:Family>
	<dwc:Genus>Lycothrissa</dwc:Genus>
	<dwc:ScientificName>Lycothrissa crocodilus (Bleeker, 1850)</dwc:ScientificName>
	<commonName xml:lang='ar'>Boefchah</commonName>
	<commonName xml:lang='my'>Nga-bah</commonName>
	<commonName xml:lang='my'>Shouk-alhin</commonName>
	<commonName xml:lang='en'>Whitehead's thryssa</commonName>
	<commonName xml:lang='et'>Whiteheadi trüssa</commonName>
	<commonName xml:lang=''>怀氏棱鳀</commonName>
	<commonName xml:lang=''>懷氏棱鯷</commonName>
	<synonym relationship='misapplied name'>Engraulis vitrirostris (non Gilchrist &amp; Thompson, 1908)</synonym>
	<synonym relationship='misapplied name'>Thrissocles purava (non Hamilton, 1822)</synonym>
	<synonym relationship='misapplied name'>Thryssa malabarica (non Bloch, 1795)</synonym>
	<synonym relationship='misapplied name'>Thryssa mystax (non Schneider, 1801)</synonym>
	<synonym relationship='misapplied name'>Thryssa purava (non Hamilton, 1822)</synonym>
	<synonym relationship='synonym'>Thryssa whiteheadi Wongratana, 1983</synonym>
	<dcterms:created>1990-10-17</dcterms:created>
	<dcterms:modified>2024-08-12</dcterms:modified>
	<reference isbn='92-5-103623-3.' url='http://www.fishbase.org/references/FBRefSummary.php?id=171&amp;speccode=603'>Coppola, S.R., W. Fischer, L. Garibaldi, N. Scialabba and K.E. Carpenter0 SPECIESDAB: Global species database for fishery purposes. User's manual. FAO Computerized Information Series (Fisheries). No. 9. Rome, FAO. 103 p. (Ref. 171)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=189&amp;speccode=603'>Whitehead, P.J.P., G.J. Nelson and T. Wongratana0 FAO Species Catalogue. Vol. 7. Clupeoid fishes of the world (Suborder Clupeoidei). An annotated and illustrated catalogue of the herrings, sardines, pilchards, sprats, shads, anchovies and wolf-herrings. FAO Fish. Synop. 125(7/2):305-579. Rome: FAO. (Ref. 189)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=6979&amp;speccode=603'>Palekar, V.C. and K.R. Karandikar0 Maturity and spawning period of &lt;i&gt;Thrissocles purava&lt;/i&gt; (Ham.) as determined by ova-diameter measurement. Proc. Indian Acad. Sci. B35(4):143-154. (Ref. 6979)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=7100&amp;speccode=603'>Khin, U.0 Fisheries in Burma. Gov't. Printing, Rangoon. 180 p. (Ref. 7100)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=11441&amp;speccode=603'>Randall, J.E.0 Coastal fishes of Oman. University of Hawaii Press, Honolulu, Hawaii. 439 p. (Ref. 11441)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=13710&amp;speccode=603'>Carpenter, K.E., F. Krupp, D.A. Jones and U. Zajonz0 FAO species identification field guide for fishery purposes. Living marine resources of Kuwait, eastern Saudi Arabia, Bahrain, Qatar, and the United Arab Emirates. FAO, Rome. 293 p. (Ref. 13710)</reference>
	<reference isbn='957-8596-59-6' url='http://www.fishbase.org/references/FBRefSummary.php?id=31517&amp;speccode=603'>Wu, H.L., K.-T. Shao and C.F. Lai (eds.)0 Latin-Chinese dictionary of fishes names. The Sueichan Press, Taiwan. 1028 p. (Ref. 31517)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=33021&amp;speccode=603'>Eschmeyer, W.N. (ed.)0 Catalog of fishes. Updated database version of November 1999. Catalog databases as made available to FishBase in November 1999. (Ref. 33021)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=40919&amp;speccode=603'>Anonymous0 Fish collection database of the Zoological Museum, University of Copenhagen. Zoological Museum, University of Copenhagen. (Ref. 40919)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=58108&amp;speccode=603'>Chinese Academy of Fishery Sciences0 Chinese aquatic germplasm resources database. http://zzzy.fishinfo.cn (Ref. 58108)</reference>
	<reference isbn='92-5-103741-8 (pbk.).' url='http://www.fishbase.org/references/FBRefSummary.php?id=68964&amp;speccode=603'>Carpenter, K.E., F. Krupp, D.A. Jones and U. Zajonz0 Living marine resources of Kuwait, eastern Saudi Arabia, Bahrain, Qatar, and the United Arab Emirates. FAO species identification field guide for fishery purposes. 324 p. Rome: FAO. (Ref. 68964)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=75549&amp;speccode=603'>FAO-FIES0 Aquatic Sciences and Fisheries Information System (ASFIS) species list. Retrieved from http://www.fao.org/fishery/collection/asfis, 29 April 2008. (Ref. 75549)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=83882&amp;speccode=603'>FAO-FIES0 Aquatic Sciences and Fisheries Information System (ASFIS) species list. Retrieved from http://www.fao.org/fishery/collection/asfis/en, March 2010. (Ref. 83882)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=90062&amp;speccode=603'>FAO-FIES0 Aquatic Sciences and Fisheries Information System (ASFIS) species list. Retrieved from http://www.fao.org/fishery/collection/asfis/en, March 2012. (Ref. 90062)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=95632&amp;speccode=603'>FAO-FIES0 Aquatic Sciences and Fisheries Information System (ASFIS) species list. Retrieved from http://www.fao.org/fishery/collection/asfis/en, April 2014. (Ref. 95632)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=101110&amp;speccode=603'>FAO-FIES0 Aquatic Sciences and Fisheries Information System (ASFIS) species list. Retrieved from http://www.fao.org/fishery/collection/asfis/en, [accessed 13/04/2015]. (Ref. 101110)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=105471&amp;speccode=603'>eElurikkus0 eElurikkus koondab ühte portaali Eesti eluslooduse andmed. http://elurikkus.ut.ee/ (Ref. 105471)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=109283&amp;speccode=603'>Coad, B.W.0 Freshwater fishes of Iraq. Pensoft Series Faunistica No. 93.  Pensoft Publishers, Moscow.  274pp. (Ref. 109283)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=115257&amp;speccode=603'>FAO-FIES0 Aquatic Sciences and Fisheries Information System (ASFIS) species list. Retrieved from http://www.fao.org/fishery/collection/asfis/en (accessed 08/06/2017). (Ref. 115257)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=115344&amp;speccode=603'>Coad, B.W.0 Fishes of the Tigris-Euphrates Basin:  a critical checklist. Syllogeus (68):1-56. (Ref. 115344)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=118272&amp;speccode=603'>FAO-FIES0 Aquatic Sciences and Fisheries Information System (ASFIS) species list. Retrieved from htttps://www.fao.org/fishery/collection/asfis/en [accessed May, 2018] (Ref. 118272)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=123974&amp;speccode=603'>Hata, H., P.N. Psomadakis, H.B. Osmany and H. Motomura0 A new species of &lt;i&gt;Thrissina&lt;/i&gt; from Pakistan (Arabian Sea), with redescription of &lt;i&gt;Thrissina whiteheadi&lt;/i&gt; (Wongratana 1983) (Clupeiformes: Engraulidae). Ichthyol. Res. 68(4):486-495. (Ref. 123974)</reference>
	<reference isbn='1849954984' url='http://www.fishbase.org/references/FBRefSummary.php?id=128868&amp;speccode=603'>Beolens, B., M. Grayson and M. Watkins0 Eponym dictionary of Fishes. 	Dunbeath, Caithness (Scotland, UK): Whittles Publishing Limited, 1300 p. (Ref. 128868)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=130435&amp;speccode=603'>IUCN0 The IUCN Red List of Threatened Species. 1964 - onwards. https://www.iucnredlist.org/ (Ref. 130435)</reference><dataObject>
	<dc:identifier>FB-pic-603-29147</dc:identifier>
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	<agent homepage='http://www.fishbase.us/collaborators/CollaboratorSummary.php?id=823' logoURL='' role='photographer'> FAO</agent>
	<dcterms:created>2001-05-15</dcterms:created>
	<dcterms:modified>2010-07-09</dcterms:modified>
	<dc:title></dc:title>
	<dc:language></dc:language>
	<license>http://creativecommons.org/licenses/by-nc/3.0/</license>
	<dc:rights>This work is licensed under a Creative Commons Attribution-Noncommercial 3.0 Unported License. Image can be freely copied and altered but may be used only for non-commercial purposes. Original author and source must be properly acknowledged</dc:rights>
	<dcterms:rightsHolder>WorldFish Center - FishBase</dcterms:rightsHolder>
	<dcterms:bibliographicCitation></dcterms:bibliographicCitation>
	<audience>General public</audience>
	<dc:source>http://www.fishbase.org/summary/SpeciesSummary.php?id=603</dc:source>
	<subject>http://rs.tdwg.org/ontology/voc/SPMInfoItems#LifeCycle</subject>
	<dc:description></dc:description>
	<mediaURL>http://www.fishbase.us/images/species/Thwhi_u0.gif</mediaURL>
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	<dc:identifier>FB-GeneralDescr-603</dc:identifier>
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	<agent homepage='' logoURL='' role='recorder'>Crispina B. Binohlan</agent>
	<dcterms:created>1990-10-17 00:00:00</dcterms:created>
	<dcterms:modified>2023-03-30 00:00:00</dcterms:modified>
	<dc:title>Biology</dc:title>
	<dc:language></dc:language>
	<license>http://creativecommons.org/licenses/by-nc/3.0/</license>
	<dc:rights></dc:rights>
	<dcterms:rightsHolder>WorldFish Center - FishBase</dcterms:rightsHolder>
	<dcterms:bibliographicCitation></dcterms:bibliographicCitation>
	<audience>General public</audience>
	<dc:source>http://www.fishbase.org/summary/SpeciesSummary.php?id=603</dc:source>
	<subject>http://rs.tdwg.org/ontology/voc/SPMInfoItems#Size</subject>
	<dc:description>Pelagic (Ref. 68964). Presumably schooling.  More data needed.  Apparently common but seldom seen in the market at Kuwait.</dc:description>
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	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=123974&amp;speccode=603'>Hata, H., P.N. Psomadakis, H.B. Osmany and H. Motomura 2021 A new species of &lt;i&gt;Thrissina&lt;/i&gt; from Pakistan (Arabian Sea), with redescription of &lt;i&gt;Thrissina whiteheadi&lt;/i&gt; (Wongratana 1983) (Clupeiformes: Engraulidae). Ichthyol. Res. 68(4):486-495. (Ref. 123974)</reference></dataObject><dataObject>
	<dc:identifier>FB-DiagnosticDescr-603</dc:identifier>
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	<agent homepage='' logoURL='' role='recorder'>Estelita Emily Capuli</agent>
	<dcterms:created>2023-03-30 00:00:00</dcterms:created>
	<dcterms:modified>2023-03-30 00:00:00</dcterms:modified>
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	<audience>General public</audience>
	<dc:source>http://www.fishbase.org/summary/SpeciesSummary.php?id=603</dc:source>
	<subject>http://rs.tdwg.org/ontology/voc/SPMInfoItems#Threats</subject>
	<dc:description>This species is distinguished by the following set of characters: upper jaw is relatively long, with its posterior tip slightly beyond opercle posterior margin, but not reaching pectoral-fin insertion; the first supramaxilla absent; teeth on both jaws are robust and enlarged; dorsum no dark lines when preserved; the dark black blotch behind upper part of gill opening is absent or indistinct in preserved specimens; branched rays of dorsal fin 9-11 (modally 10), anal fin usually 39-42 (rarely 43, modally 40) and pectoral fin 11-12 (modally 12); transverse scales 10-11 (11); scale rows in longitudinal series 40-42 (42); gill rakers on first gill arch, in upper series 12–15 (13), lower series 18-21 (19), total 30-34 (rarely 36, modally 32); gill rakers on second gill arch, in upper series 10-13 (modally 12), lower series 18-21 (19), total 29-34 (31); gill rakers on third gill arch, in upper series 8-10 (8), lower series 10-13 (11), total 19-23 (19); gill rakers on fourth gill arch, in upper series 6-8 (7), lower series 8-11 (10), total 14-19 (17) in total; snout is short, 3.1-3.6% SL pelvic fin short, 7.7-9.0% SL; mandible is short, 15.4-17.2% SL; pelvic-fin insertion to anal-fin origin 19.1-24.2% SL (Ref. 123974).</dc:description>
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	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=123974&amp;speccode=603'>Hata, H., P.N. Psomadakis, H.B. Osmany and H. Motomura 2021 A new species of &lt;i&gt;Thrissina&lt;/i&gt; from Pakistan (Arabian Sea), with redescription of &lt;i&gt;Thrissina whiteheadi&lt;/i&gt; (Wongratana 1983) (Clupeiformes: Engraulidae). Ichthyol. Res. 68(4):486-495. (Ref. 123974)</reference></dataObject><dataObject>
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	<dcterms:created>1990-10-18 00:00:00</dcterms:created>
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	<subject>http://rs.tdwg.org/ontology/voc/SPMInfoItems#Habitat</subject>
	<dc:description>Western Indian Ocean: only in the Persian Gulf, but may reach into the Gulf of Oman.</dc:description>
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	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=189&amp;speccode=603'>Whitehead, P.J.P., G.J. Nelson and T. Wongratana 1988 FAO Species Catalogue. Vol. 7. Clupeoid fishes of the world (Suborder Clupeoidei). An annotated and illustrated catalogue of the herrings, sardines, pilchards, sprats, shads, anchovies and wolf-herrings. FAO Fish. Synop. 125(7/2):305-579. Rome: FAO. (Ref. 189)</reference></dataObject><dataObject>
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	<dc:title>Importance</dc:title>
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	<dcterms:rightsHolder>John E. Randall</dcterms:rightsHolder>
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	<subject></subject>
	<dc:description>fisheries: subsistence fisheries</dc:description>
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	<reference isbn='92-5-103623-3.' url='http://www.fishbase.org/references/FBRefSummary.php?id=171&amp;speccode=603'>Coppola, S.R., W. Fischer, L. Garibaldi, N. Scialabba and K.E. Carpenter 1994 SPECIESDAB: Global species database for fishery purposes. User's manual. FAO Computerized Information Series (Fisheries). No. 9. Rome, FAO. 103 p. (Ref. 171)</reference></dataObject><dataObject>
	<dc:identifier>FB-Size-603</dc:identifier>
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	<dc:title>Max. size</dc:title>
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	<subject>http://rs.tdwg.org/ontology/voc/SPMInfoItems#DiagnosticDescription</subject>
	<dc:description>20.5 cm SL (male/unsexed; (Ref. 123974))</dc:description>
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	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=123974&amp;speccode=603'>Hata, H., P.N. Psomadakis, H.B. Osmany and H. Motomura 2021 A new species of &lt;i&gt;Thrissina&lt;/i&gt; from Pakistan (Arabian Sea), with redescription of &lt;i&gt;Thrissina whiteheadi&lt;/i&gt; (Wongratana 1983) (Clupeiformes: Engraulidae). Ichthyol. Res. 68(4):486-495. (Ref. 123974)</reference></dataObject><dataObject>
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	<subject>http://rs.tdwg.org/ontology/voc/SPMInfoItems#Morphology</subject>
	<dc:description>Least Concern (LC) , IUCN Grouper and Wrasse Specialist Group</dc:description>
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	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=57073'>IUCN 2006 2006 IUCN red list of threatened species. www.iucnredlist.org. Downloaded July 2006.</reference></dataObject><dataObject>
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	<dc:title>Environment</dc:title>
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	<audience>General public</audience>
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	<subject>http://rs.tdwg.org/ontology/voc/SPMInfoItems#Distribution</subject>
	<dc:description>pelagic-neritic; brackish; marine</dc:description>
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