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<taxon>
	<dc:identifier>FB-5837</dc:identifier>
	<dc:source>http://www.fishbase.org/summary/SpeciesSummary.php?id=5837</dc:source>
	<dwc:Kingdom>Animalia</dwc:Kingdom>
	<dwc:Phylum>Chordata</dwc:Phylum>
	<dwc:Class>Teleostei</dwc:Class>
	<dwc:Order>Perciformes/Serranoidei</dwc:Order>
	<dwc:Family>Epinephelidae</dwc:Family>
	<dwc:Genus>Epinephelus</dwc:Genus>
	<dwc:ScientificName>Epinephelus spilotoceps Schultz, 1953</dwc:ScientificName>
	<commonName xml:lang='pt'>Garoupa de quatro selas</commonName>
	<commonName xml:lang='af'>Foursaddle rockcod</commonName>
	<commonName xml:lang='af'>Viersaal-klipkabeljou</commonName>
	<commonName xml:lang='de'>Zackenbarsch</commonName>
	<commonName xml:lang='en'>Foursaddle grouper</commonName>
	<commonName xml:lang='en'>Four-saddle grouper</commonName>
	<commonName xml:lang='en'>Spotty cod</commonName>
	<commonName xml:lang='fj'>senicauca</commonName>
	<commonName xml:lang='fr'>Mérou quatre selles</commonName>
	<commonName xml:lang='fr'>Vieille fond</commonName>
	<commonName xml:lang='he'>Lokos</commonName>
	<commonName xml:lang='ja'>Yotsugurabata</commonName>
	<commonName xml:lang='mh'>Mur in ae en</commonName>
	<commonName xml:lang='ms'>Kerapu cicak</commonName>
	<commonName xml:lang='es'>Mero cuatro monturas</commonName>
	<commonName xml:lang=''>Asdhaan faana</commonName>
	<commonName xml:lang=''>Gaily mwera</commonName>
	<commonName xml:lang=''>Macabit</commonName>
	<commonName xml:lang=''>吻斑石斑魚</commonName>
	<commonName xml:lang=''>吻斑石斑鱼</commonName>
	<commonName xml:lang=''>花頭梅</commonName>
	<synonym relationship='synonym'>Epinephelus salonotus Smith &amp; Smith, 1963</synonym>
	<synonym relationship='synonym'>Epinephelus spiloteceps Schultz, 1953</synonym>
	<dcterms:created>1992-06-15</dcterms:created>
	<dcterms:modified>2022-02-09</dcterms:modified>
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	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=118272&amp;speccode=5837'>FAO-FIES0 Aquatic Sciences and Fisheries Information System (ASFIS) species list. Retrieved from htttps://www.fao.org/fishery/collection/asfis/en [accessed May, 2018] (Ref. 118272)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=118580&amp;speccode=5837'>SCRFA, Science and Conservation of Fish Aggregations0 SCRFA database. Web online database. https://www.scrfa.org/database/: SCRFA. [2013-onwards]. (Ref. 118580)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=118793&amp;speccode=5837'>Allen, G.R. and M.V. Erdmann0 Coral reef fishes of Timor-Leste. p. 33-82.  In G.R. Allen and M.V. Erdmann (eds).  A rapid biological assessment of Timor-Leste. Conservation International. (Ref. 118793)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=122752&amp;speccode=5837'>University of Western Australia0 Benthic  BRUVS data 2019. Video BRUVS from Jessica Meeuwig. (Ref. 122752)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=124899&amp;speccode=5837'>van der Elst, Rudy, Simon Chater and Dennis King0 Fishes from Bassas Da India Atoll in the Mozambique Channel. Atoll Research Bulletin No. 629. (Ref. 124899)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=125599&amp;speccode=5837'>Matthews, T., D. Ochavillo, S. Felise, T. Letalie, E. Letuane, E. Schuster, A. Soonaolo, S. Tofaeono, A. Tua and F. Tuilagi0 Length-weight relationships for 71 reef and bottomfish species from Tutuila and Aunu'u, American Samoa. Pacific Islands Fish. Sci. Cent. Admin. Rep. H-19-03, 9 p. (Ref. 125599)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=137577&amp;speccode=5837'>Price, S.A., S.T. Friedman, K.A. Corn, O. Larouche, K. Brockelsby, A.J. Lee, M. Nagaraj, N.G. Bertrand, M. Danao, M.C. Coyne, J.R. Estrada, R. Friedman, E. Hoeft, M. Iwan, D. Gross, J.H. Kao, B. Landry, M.J. Linares, C. McGlinn and …0 FishShapes v1: Functionally relevant measurements of teleost shape and size on three dimensions. Ecology 103(e3829):1-2. (Ref. 137577)</reference><dataObject>
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	<agent homepage='http://www.fishbase.us/collaborators/CollaboratorSummary.php?id=50' logoURL='' role='photographer'>John E. Randall</agent>
	<dcterms:created>1996-03-04</dcterms:created>
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	<dc:rights>This work is licensed under a Creative Commons Attribution-Noncommercial 3.0 Unported License. Image can be freely copied and altered but may be used only for non-commercial purposes. Original author and source must be properly acknowledged</dc:rights>
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	<dc:description>outer reef; top: 4.4 SL, 5.6 TL; depth range: 1-1.5 m</dc:description>
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	<mediaURL>http://www.fishbase.us/images/species/Epspi_u2.jpg</mediaURL>
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	<dc:description></dc:description>
	<mediaURL>http://www.fishbase.us/images/species/Epspi_u3.jpg</mediaURL>
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	<dc:identifier>FB-pic-5837-9625</dc:identifier>
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	<agent homepage='http://www.fishbase.us/collaborators/CollaboratorSummary.php?id=50' logoURL='' role='photographer'>John E. Randall</agent>
	<dcterms:created>1996-03-04</dcterms:created>
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	<dc:rights>This work is licensed under a Creative Commons Attribution-Noncommercial 3.0 Unported License. Image can be freely copied and altered but may be used only for non-commercial purposes. Original author and source must be properly acknowledged</dc:rights>
	<dcterms:rightsHolder>WorldFish Center - FishBase</dcterms:rightsHolder>
	<dcterms:bibliographicCitation></dcterms:bibliographicCitation>
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	<dc:source>http://www.fishbase.org/summary/SpeciesSummary.php?id=5837</dc:source>
	<subject>http://rs.tdwg.org/ontology/voc/SPMInfoItems#LifeCycle</subject>
	<dc:description></dc:description>
	<mediaURL>http://www.fishbase.us/images/species/Epspi_u4.jpg</mediaURL>
	<thumbnailURL>http://www.fishbase.us/images/thumbnails/jpg/tn_Epspi_u4.jpg</thumbnailURL>
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	<dc:identifier>FB-GeneralDescr-5837</dc:identifier>
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	<agent homepage='' logoURL='' role='recorder'>Crispina B. Binohlan</agent>
	<dcterms:created>1992-06-15 00:00:00</dcterms:created>
	<dcterms:modified>2022-02-09 00:00:00</dcterms:modified>
	<dc:title>Biology</dc:title>
	<dc:language></dc:language>
	<license>http://creativecommons.org/licenses/by-nc/3.0/</license>
	<dc:rights></dc:rights>
	<dcterms:rightsHolder>WorldFish Center - FishBase</dcterms:rightsHolder>
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	<audience>General public</audience>
	<dc:source>http://www.fishbase.org/summary/SpeciesSummary.php?id=5837</dc:source>
	<subject>http://rs.tdwg.org/ontology/voc/SPMInfoItems#TrophicStrategy</subject>
	<dc:description>Occurs in lagoon patch reefs, the upper slopes of channels, and reef margins (Ref. 1602).  Carnivorous (Ref. 81697).  Nothing has been published on its biology. Solitary (Ref 90102).</dc:description>
	<mediaURL></mediaURL>
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	<location></location>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=5222&amp;speccode=5837'>Heemstra, P.C. and J.E. Randall 1993 FAO Species Catalogue. Vol. 16. Groupers of the world (family Serranidae, subfamily Epinephelinae). An annotated and illustrated catalogue of the grouper, rockcod, hind, coral grouper and lyretail species known to date. Rome: FAO. FAO Fish. Synop. 125(16):382 p. (Ref. 5222)</reference></dataObject><dataObject>
	<dc:identifier>FB-DiagnosticDescr-5837</dc:identifier>
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	<agent homepage='https://sites.google.com/site/itsaboutfish/home#' logoURL='' role='recorder'>Rodolfo B. Reyes</agent>
	<dcterms:created>1994-06-14 00:00:00</dcterms:created>
	<dcterms:modified>2013-03-11 00:00:00</dcterms:modified>
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	<audience>General public</audience>
	<dc:source>http://www.fishbase.org/summary/SpeciesSummary.php?id=5837</dc:source>
	<subject>http://rs.tdwg.org/ontology/voc/SPMInfoItems#Size</subject>
	<dc:description>Preopercle rounded, with a shallow notch; the serrae are enlarged but covered with skin; upper edge of the operculum almost straight.  The head and body mostly covered with close-set dark brown, olive, or reddish brown spots.  Those on the dorsolateral parts of the head, body, and on median fins polygonal and close-set, with the interspaces forming a white reticulum.  Those on the ventral portions are more rounded, more separated and often more reddish.  Three dark blotches are often visible at the base of the dorsal fin and one forming a saddle blotch on the caudal peduncle.  These blotches differentiates it from E. merra and E. quoyanus (Ref. 5222).  Small conspicuous dark spots on snout and sometimes between or below the eyes (Ref. 37816).</dc:description>
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	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=5222&amp;speccode=5837'>Heemstra, P.C. and J.E. Randall 1993 FAO Species Catalogue. Vol. 16. Groupers of the world (family Serranidae, subfamily Epinephelinae). An annotated and illustrated catalogue of the grouper, rockcod, hind, coral grouper and lyretail species known to date. Rome: FAO. FAO Fish. Synop. 125(16):382 p. (Ref. 5222)</reference></dataObject><dataObject>
	<dc:identifier>FB-Morphology-5837</dc:identifier>
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	<agent homepage='https://sites.google.com/site/itsaboutfish/home#' logoURL='' role='recorder'>Rodolfo B. Reyes</agent>
	<dcterms:created>1994-06-14 00:00:00</dcterms:created>
	<dcterms:modified>2013-03-11 00:00:00</dcterms:modified>
	<dc:title></dc:title>
	<dc:language></dc:language>
	<license>http://creativecommons.org/licenses/by-nc/3.0/</license>
	<dc:rights></dc:rights>
	<dcterms:rightsHolder>WorldFish Center - FishBase</dcterms:rightsHolder>
	<dcterms:bibliographicCitation></dcterms:bibliographicCitation>
	<audience>General public</audience>
	<dc:source>http://www.fishbase.org/summary/SpeciesSummary.php?id=5837</dc:source>
	<subject>http://rs.tdwg.org/ontology/voc/SPMInfoItems#Threats</subject>
	<dc:description>Dorsal spines (total): 11; Dorsal soft rays (total): 14 - 16;  Analspines: 3; Analsoft rays: 8</dc:description>
	<mediaURL></mediaURL>
	<thumbnailURL></thumbnailURL>
	<location></location>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=5222&amp;speccode=5837'>Heemstra, P.C. and J.E. Randall 1993 FAO Species Catalogue. Vol. 16. Groupers of the world (family Serranidae, subfamily Epinephelinae). An annotated and illustrated catalogue of the grouper, rockcod, hind, coral grouper and lyretail species known to date. Rome: FAO. FAO Fish. Synop. 125(16):382 p. (Ref. 5222)</reference></dataObject><dataObject>
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	<agent homepage='' logoURL='' role='recorder'>Crispina B. Binohlan</agent>
	<dcterms:created>1992-06-15 00:00:00</dcterms:created>
	<dcterms:modified>2012-05-07 00:00:00</dcterms:modified>
	<dc:title></dc:title>
	<dc:language></dc:language>
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	<dcterms:bibliographicCitation></dcterms:bibliographicCitation>
	<audience>General public</audience>
	<dc:source>http://www.fishbase.org/summary/SpeciesSummary.php?id=5837</dc:source>
	<subject>http://rs.tdwg.org/ontology/voc/SPMInfoItems#Habitat</subject>
	<dc:description>Indo-West-Pacific:  East coast of Africa to the Line Islands.  Except for its occurrence along the African coast (Zanzibar, Tanzania to Ponta Zavora, Mozambique) it seems to be primarily an insular species, occurring at most (probably all) of the islands of tropical Indian and west-central Pacific.  Not known from the Red Sea, Persian Gulf, Sri Lanka, Philippines, Taiwan, Japan, or Australian waters (however, it is found at Rowley Shoals off Western Australia).</dc:description>
	<mediaURL></mediaURL>
	<thumbnailURL></thumbnailURL>
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	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=5222&amp;speccode=5837'>Heemstra, P.C. and J.E. Randall 1993 FAO Species Catalogue. Vol. 16. Groupers of the world (family Serranidae, subfamily Epinephelinae). An annotated and illustrated catalogue of the grouper, rockcod, hind, coral grouper and lyretail species known to date. Rome: FAO. FAO Fish. Synop. 125(16):382 p. (Ref. 5222)</reference></dataObject><dataObject>
	<dc:identifier>FB-Uses-5837</dc:identifier>
	<dataType>http://purl.org/dc/dcmitype/Text</dataType>
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	<agent homepage='' logoURL='' role='recorder'>Crispina B. Binohlan</agent>
	<dcterms:created>1992-06-15 00:00:00</dcterms:created>
	<dcterms:modified>2022-02-09 00:00:00</dcterms:modified>
	<dc:title>Importance</dc:title>
	<dc:language></dc:language>
	<license>http://creativecommons.org/licenses/by-nc/3.0/</license>
	<dc:rights></dc:rights>
	<dcterms:rightsHolder>WorldFish Center - FishBase</dcterms:rightsHolder>
	<dcterms:bibliographicCitation></dcterms:bibliographicCitation>
	<audience>General public</audience>
	<dc:source>http://www.fishbase.org/summary/SpeciesSummary.php?id=5837</dc:source>
	<subject>http://rs.tdwg.org/ontology/voc/SPMInfoItems#DiagnosticDescription</subject>
	<dc:description>fisheries: commercial; gamefish: yes</dc:description>
	<mediaURL></mediaURL>
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	<location></location>
	<reference isbn='92-5-103623-3.' url='http://www.fishbase.org/references/FBRefSummary.php?id=171&amp;speccode=5837'>Coppola, S.R., W. Fischer, L. Garibaldi, N. Scialabba and K.E. Carpenter 1994 SPECIESDAB: Global species database for fishery purposes. User's manual. FAO Computerized Information Series (Fisheries). No. 9. Rome, FAO. 103 p. (Ref. 171)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=30874&amp;speccode=5837'>Allen, G.R. and R.C. Steene 1988 Fishes of Christmas Island Indian Ocean. Christmas Island Natural History Association, Christmas Island, Indian Ocean, 6798, Australia. 197 p. (Ref. 30874)</reference></dataObject><dataObject>
	<dc:identifier>FB-TrophicStrategy-5837</dc:identifier>
	<dataType>http://purl.org/dc/dcmitype/Text</dataType>
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	<agent homepage='' logoURL='' role='recorder'>Auda Kareen Ortañez</agent>
	<dcterms:created>2007-08-09 00:00:00</dcterms:created>
	<dcterms:modified>2009-09-08 00:00:00</dcterms:modified>
	<dc:title></dc:title>
	<dc:language></dc:language>
	<license>http://creativecommons.org/licenses/by-nc/3.0/</license>
	<dc:rights></dc:rights>
	<dcterms:rightsHolder>WorldFish Center - FishBase</dcterms:rightsHolder>
	<dcterms:bibliographicCitation></dcterms:bibliographicCitation>
	<audience>General public</audience>
	<dc:source>spilotoceps</dc:source>
	<subject>http://rs.tdwg.org/ontology/voc/SPMInfoItems#Morphology</subject>
	<dc:description>Found inshore (Ref. 75154).</dc:description>
	<mediaURL></mediaURL>
	<thumbnailURL></thumbnailURL>
	<location></location>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=58534&amp;speccode=5837'>Nguyen, N.T. and V.Q. Nguyen 2006 Biodiversity and living resources of the coral reef fishes in Vietnam marine waters. Science and Technology Publishing House, Hanoi. (Ref. 58534)</reference></dataObject><dataObject>
	<dc:identifier>FB-Size-5837</dc:identifier>
	<dataType>http://purl.org/dc/dcmitype/Text</dataType>
	<mimeType>text/html</mimeType>
	<agent homepage='' logoURL='' role='recorder'>Crispina B. Binohlan</agent>
	<dcterms:created>1992-06-15 00:00:00</dcterms:created>
	<dcterms:modified>2022-02-09 00:00:00</dcterms:modified>
	<dc:title>Max. size</dc:title>
	<dc:language></dc:language>
	<license>http://creativecommons.org/licenses/by-nc/3.0/</license>
	<dc:rights></dc:rights>
	<dcterms:rightsHolder>WorldFish Center - FishBase</dcterms:rightsHolder>
	<dcterms:bibliographicCitation></dcterms:bibliographicCitation>
	<audience>General public</audience>
	<dc:source>http://www.fishbase.org/summary/SpeciesSummary.php?id=5837</dc:source>
	<subject>http://rs.tdwg.org/ontology/voc/SPMInfoItems#Distribution</subject>
	<dc:description>47.5 cm TL (male/unsexed; (Ref. 125599)); max. published weight: 1,543 g (Ref. 125599)</dc:description>
	<mediaURL></mediaURL>
	<thumbnailURL></thumbnailURL>
	<location></location>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=125599&amp;speccode=5837'>Matthews, T., D. Ochavillo, S. Felise, T. Letalie, E. Letuane, E. Schuster, A. Soonaolo, S. Tofaeono, A. Tua and F. Tuilagi 2019 Length-weight relationships for 71 reef and bottomfish species from Tutuila and Aunu'u, American Samoa. Pacific Islands Fish. Sci. Cent. Admin. Rep. H-19-03, 9 p. (Ref. 125599)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=125599&amp;speccode=5837'>Matthews, T., D. Ochavillo, S. Felise, T. Letalie, E. Letuane, E. Schuster, A. Soonaolo, S. Tofaeono, A. Tua and F. Tuilagi 2019 Length-weight relationships for 71 reef and bottomfish species from Tutuila and Aunu'u, American Samoa. Pacific Islands Fish. Sci. Cent. Admin. Rep. H-19-03, 9 p. (Ref. 125599)</reference></dataObject><dataObject>
	<dc:identifier>FB-Threats-2</dc:identifier>
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	<agent homepage='' logoURL='' role='recorder'>Belen Acosta</agent>
	<dcterms:created>1990-10-18 00:00:00</dcterms:created>
	<dcterms:modified>2024-03-07 00:00:00</dcterms:modified>
	<dc:title></dc:title>
	<dc:language></dc:language>
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	<dcterms:rightsHolder>WorldFish Center - FishBase</dcterms:rightsHolder>
	<dcterms:bibliographicCitation></dcterms:bibliographicCitation>
	<audience>General public</audience>
	<dc:source>http://www.fishbase.org/summary/SpeciesSummary.php?id=2</dc:source>
	<subject>http://rs.tdwg.org/ontology/voc/SPMInfoItems#Uses</subject>
	<dc:description>Least Concern (LC) , IUCN Grouper and Wrasse Specialist Group</dc:description>
	<mediaURL></mediaURL>
	<thumbnailURL></thumbnailURL>
	<location></location>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=57073'>IUCN 2006 2006 IUCN red list of threatened species. www.iucnredlist.org. Downloaded July 2006.</reference></dataObject><dataObject>
	<dc:identifier>FB-Habitat-5837</dc:identifier>
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	<agent homepage='' logoURL='' role='recorder'>Crispina B. Binohlan</agent>
	<dcterms:created>1992-06-15 00:00:00</dcterms:created>
	<dcterms:modified>2022-02-09 00:00:00</dcterms:modified>
	<dc:title>Environment</dc:title>
	<dc:language></dc:language>
	<license>http://creativecommons.org/licenses/by-nc/3.0/</license>
	<dc:rights></dc:rights>
	<dcterms:rightsHolder>WorldFish Center - FishBase</dcterms:rightsHolder>
	<dcterms:bibliographicCitation></dcterms:bibliographicCitation>
	<audience>General public</audience>
	<dc:source>http://www.fishbase.org/summary/SpeciesSummary.php?id=5837</dc:source>
	<subject>http://rs.tdwg.org/ontology/voc/SPMInfoItems#Size</subject>
	<dc:description>reef-associated; marine; depth range 0 - 30 m (Ref. 9710)</dc:description>
	<mediaURL></mediaURL>
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	<reference isbn='0-00-219974-2 (pbk.).' url='http://www.fishbase.org/references/FBRefSummary.php?id=9710&amp;speccode=5837'>Lieske, E. and R. Myers 1994 Collins Pocket Guide. Coral reef fishes. Indo-Pacific &amp; Caribbean including the Red Sea. Haper Collins Publishers, 400 p. (Ref. 9710)</reference></dataObject>
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