FB-5580 http://www.fishbase.org/summary/SpeciesSummary.php?id=5580 Animalia Chordata Teleostei Acanthuriformes Chaetodontidae Chaetodon Chaetodon ulietensis Cuvier, 1831 Alibangbang Dobbeltsadlet fanefisk Doublesaddle butterflyfish Double-saddle butterflyfish Ovalspot butterflyfish Pacific double-saddled butterflyfish Pacific doublesaddled coralfish Pacific doublesaddle butterflyfish Pacific double-saddle butterflyfish Sickle butterflyfish daliga ragutu Papillon à deux selles du Pacifique Papillon à selle Paraharaha Sudare-chôchô-uo Bagang Bagang dwipelana Kitang Tifitifi-gutu'uli Paru-paro Alibang-bang Arulole Kapiged 乌利蝴蝶鱼 烏利蝴蝶魚 雙印蝶 鞍斑蝴蝶魚 鞍斑蝴蝶鱼 鞍斑蝶 Chaetodon aurora De Vis, 1884 Chaetodon falcula (non Bloch, 1795) Chaetodon ulientensis Cuvier, 1831 Chaetodon ulietensis confluens Ahl, 1923 Rabdophorus oxychaetodon ulietensis (Cuvier, 1831) 1992-06-05 2023-06-14 Breder, C.M. and D.E. Rosen0 Modes of reproduction in fishes. T.F.H. Publications, Neptune City, New Jersey. 941 p. (Ref. 205) Thresher, R.E.0 Reproduction in reef fishes. T.F.H. Publications, Inc. Ltd., Neptune City, New Jersey. 399 p. (Ref. 240) Herre, A.W.C.T.0 Check list of Philippine fishes. Res. Rep. U.S. Fish Wild. Serv. (20):1-977. (Ref. 280) Rau, N. and A. Rau0 Commercial marine fishes of the Central Philippines (bony fish). German Agency for Technical Cooperation, Germany. 623 pp. (Ref. 393) Masuda, H., K. Amaoka, C. Araga, T. Uyeno and T. Yoshino0 The fishes of the Japanese Archipelago. Vol. 1. Tokai University Press, Tokyo, Japan. 437 p. (text). (Ref. 559) Wass, R.C.0 An annotated checklist of the fishes of Samoa. Natl. Ocean. Atmos. Adminis. Tech. Rept., Natl. Mar. Fish. Serv., Spec. Sci. Rept. Fish. (781). (Ref. 592) Myers, R.F.0 Micronesian reef fishes. Second Ed. Coral Graphics, Barrigada, Guam. 298 p. (Ref. 1602) Randall, J.E., G.R. Allen and R.C. Steene0 Fishes of the Great Barrier Reef and Coral Sea. University of Hawaii Press, Honolulu, Hawaii. 506 p. (Ref. 2334) Herre, A.W.C.T. and A.F. Umali0 English and local common names of Philippine fishes. U. S. Dept. of Interior and Fish and Wildl. Serv. Circular No. 14, U. S. Gov't Printing Office, Washington. 128 p. (Ref. 2857) Hureau, J.-C.0 La base de données GICIM: Gestion informatisée des collections ichthyologiques du Muséum. p. 225-227. In Atlas Préliminaire des Poissons d'Eau Douce de France. Conseil Supérieur de la Pêche, Ministère de l'Environnement, CEMAGREF et Muséum national d'Histoire naturelle, Paris. (Ref. 4517) Burgess, W.E.0 Butterflyfishes of the world. A monograph of the Family Chaetodontidae. T.F.H. Publications, Neptune City, New Jersey. (Ref. 4855) Steene, R.C.0 Butterfly and angelfishes of the world. A.H. & A.W. Reed Pty Ltd., Australia. vol. 1. 144 p. (Ref. 4859) Edwards, A.J. and A.D. Shepherd0 Environmental implications of aquarium-fish collection in the Maldives, with proposals for regulation. Environ. Conserv. 19:61-72. (Ref. 4907) McManus, J.W., C.L. Nañola Jr., R.B. Reyes Jr. and K.N. Kesner0 Resource ecology of the Bolinao coral reef system. ICLARM Stud. Rev. 22:117 p. (Ref. 5399) Mohsin, A.K.M., M.A. Ambak and M.N.A. Salam0 Malay, English, and scientific names of the fishes of Malaysia. Occas. Publ. Fac. Fish. Mar. Sci. Univ. Pertanian Malays. 11:226 p. (Ref. 5756) Kailola, P.J.0 The fishes of Papua New Guinea: a revised and annotated checklist. Vol. II Scorpaenidae to Callionymidae. Research Bulletin No. 41, Research Section, Dept. of Fisheries and Marine Resources, Papua New Guinea. (Ref. 6192) Kuiter, R.H.0 Tropical reef-fishes of the western Pacific Indonesia and adjacent waters. Gramedia Pustaka Utama, Jakarta. 314 p. (Ref. 8631) Francis, M.P.0 Checklist of the coastal fishes of Lord Howe, Norfolk, and Kermadec Islands, Southwest Pacific Ocean. Pac. Sci. 47(2):136-170. (Ref. 8879) Masuda, H. and G.R. Allen0 Meeresfische der Welt - Groß-Indopazifische Region. Tetra Verlag, Herrenteich, Melle. 528 p. (Ref. 9137) Lieske, E. and R. Myers0 Collins Pocket Guide. Coral reef fishes. Indo-Pacific & Caribbean including the Red Sea. Haper Collins Publishers, 400 p. (Ref. 9710) Randall, J.E. and C. Anderson0 Annotated checklist of the epipelagic and shore fishes of the Maldives Islands. Ichthyol. Bull. of the J.L.B. Smith Inst. of Ichthyol. (59):1-47. (Ref. 11303) Bauchot, R., J.M. Ridet and M.L. Bauchot0 The brain organization of butterflyfishes. Environ. Biol. Fishes 25(1-3):205-219. (Ref. 11740) Kulbicki, M., J.E. Randall and J. Rivaton0 Checklist of the fishes of the Chesterfield Islands (Coral Sea). Micronesica 27(1/2):1-43. (Ref. 11897) Kulbicki, M. and J.T. Williams0 Checklist of the shorefishes of Ouvea Atoll, New Caledonia. Atoll Res. Bull. 444:26 p. (Ref. 13236) Randall, J.E., H. Ida, K. Kato, R.L. Pyle and J.L. Earle0 Annotated checklist of inshore fishes of the Ogasawara Islands. Nat. Sci. Mus. Monogr. (11):1-74. (Ref. 26066) Foale, S.0 What's in a name? An analysis of the West Nggela (Solomon Islands) fish taxonomy. SPC Traditional Marine Resource Management and Knowledge Information Bulletin No. 9:3-19. (Ref. 26367) Mohsin, A.K.M. and M.A. Ambak0 Marine fishes and fisheries of Malaysia and neighbouring countries. University of Pertanian Malaysia Press, Serdang, Malaysia. 744 p. (Ref. 27550) Randall, J.E.0 Randall's tank photos. Collection of 10,000 large-format photos (slides) of dead fishes. Unpublished. (Ref. 28618) Allen, G.R. and P.L. Munday0 Kimbe Bay rapid ecological assessment: the coral reefs of Kimbe Bay (West New Britain, Papua New Guinea), Volume 3: Fish diversity of Kimbe Bay. The Nature Conservancy, South Pacific program Office, Auckland, New Zealand. 107 p. (Ref. 28888) Ganaden, S.R. and F. Lavapie-Gonzales0 Common and local names of marine fishes of the Philippines. Bureau of Fisheries and Aquatic Resources, Philippines. 385 p. (Ref. 31411) Wu, H.L., K.-T. Shao and C.F. Lai (eds.)0 Latin-Chinese dictionary of fishes names. The Sueichan Press, Taiwan. 1028 p. (Ref. 31517) Anonymous0 Fish collection database of the Bernice P. Bishop Museum (BPBM). Bishop Museum, 1525 Bernice Street, Honolulu, Hawai`I, 96817-0916 USA. (Ref. 31667) McCormack, G.0 Cook Islands biodiversity and natural heritage database. Sent by Gerald McCormack as RTF document in May 2000 for use in FishBase. (Ref. 34027) Randall, J.E. and K.K.P. Lim (eds.)0 A checklist of the fishes of the South China Sea. Raffles Bull. Zool. Suppl. (8):569-667. (Ref. 36648) Myers, R.F.0 Micronesian reef fishes: a comprehensive guide to the coral reef fishes of Micronesia, 3rd revised and expanded edition. Coral Graphics, Barrigada, Guam. 330 p. (Ref. 37816) Anonymous0 Fish collection database of the National Museum of Natural History (Smithsonian Institution). Smithsonian Institution-Division of Fishes. (Ref. 38732) Findley, J.S. and M.T. Findley0 Global, regional, and local patterns in species richness and abundance of butterflyfishes. Ecological Monographs. 7(1), pp. 69-91. (Ref. 40521) Eschmeyer, W.N. (ed.)0 Catalog of fishes. Updated database version of December 2001. Catalog databases as made available to FishBase in December 2001. (Ref. 40966) Anonymous0 Fish collection database of the American Museum of Natural History. American Museum of Natural History, Central Park West , NY 10024-5192, USA. (Ref. 41414) Allen, G.R.0 Reef fishes of Milne Bay Province, Papua New Guinea. In T. Werner and G. Allen (eds). A rapid biodiversity assessment of the coral reefs of Milne Bay Province, Papua New Guinea. RAP Working Papers 11, Conservation International, Washington, D.C. (Ref. 41464) Randall, J.E.0 Report on fish collections from the Pitcairn Islands. Atoll Res. Bull. (461):1-36. (Ref. 41650) White, A. and H. Calumpong0 Saving Tubbataha Reef: Earthwatch expedition to the Philippines '92. Silliman J. 36(2):77-105. (Ref. 42484) Harborne, A.R., A.B. Gill, P.S. Raines and J.M. Ridley0 Danjugan Island marine reserve. Summary report to the Philippine reef and Rainforest Conservation Foundation. (Ref. 42831) Nakabo, T.0 Fishes of Japan with pictorial keys to the species, English edition II. Tokai University Press, Japan, pp 867-1749. (Ref. 43239) Dioneda, R.R., L.R. Pura, Q.P. Sia III and L.O. Basmayor0 A checklist of fishes and invertebrates caught and observed in Lagonoy Gulf. <i>In</I> G. Silvestre, C. Luna, V. Soliman and L. Garces (eds.) Resource and ecological assessment of Lagonoy Gulf, Philippines: Terminal Report Vol.2 Technical Monograph. ICLARM Tech Rep. 000,000 p. (Ref. 43283) Anonymous0 Fish collection of the University of the Philippines in the Visayas Museum. UPV Museum. (Ref. 43309) Silvestre, G.T., L.R. Garces and C.Z. Luna0 Resource and ecological assessment of Lagonoy Gulf, Philippines: Terminal Report Vol.1 Final Report. In G. Silvestre, C. Luna, V. Soliman and L. Garces (eds.) Resource and ecological assessment of Lagonoy Gulf, Philippines. ICLARM Tech. Rep. 000, 000 p. (Ref. 45227) Nañola, C.L. and J.P. Cabansag0 Coral reef fish species and abundance of Lagonoy Gulf. In G. Silvestre, C. Luna, V. Soliman and L. Garces (eds.) Resource and ecological assessment of Lagonoy Gulf, Philippines: Terminal Report Vol.2 Technical Monograph. ICLARM Tech Rep. 000,000 p. (Ref. 45291) Garces, L.R., Q.P. Sia, M.J.M. Vega, J. Cabansag and V. Hilomen0 Species composition and abundance of reef fish communities. p. 000-000. In G. Silvestre, C. Luna and J. Padilla (eds.) Multidisciplinary assessment of the fisheries in San Miguel Bay, Philippines (1992-1993). ICLARM Technical Report 47. International Center for Living Aquatic Resources Management, Makati, Philippines. (Ref. 45334) Dantis, A.L. and P.M. Aliño (comps.)0 Checklist of Philippine reef fishes. p. 208-226. In Aliño, P.M.,E.F.B.Miclat,C.L.Nañola Jr.,H.A.Roa-Quiaoit and R.T.Campos(eds.) Atlas of Philippine coral reefs.Philippine Coral Reef Information (Philreefs). Goodwill Trading Co.,Inc.(Goodwill Bookstore),Quezon City,Philippines. xvi + 264p. (Ref. 45419) Calumpong, H.P., L.J. Raymundo and E.P. Solis-Duran (eds.)0 Resource and ecological assessment of of Sogod Bay, Leyte, Philippines - Final Report Vol.1 Fisheries Assessment. Siliman University Marine Laboratory. (Ref. 46392) Tettelbach, S., J. Carroll and H. Reisman0 Fishes of Vanuatu. Report of the Tropical Marine Biology group, Southampton College of Long Island University, Southampton, New York. (Ref. 46904) Anonymous0 Fish collection of the Royal Ontario Museum. Royal Ontario Museum. (Ref. 47438) Allen, G.R. and M. Adrim0 Coral reef fishes of Indonesia. Zool. Stud. 42(1):1-72. (Ref. 47567) Huang, Z.0 Marine species and their distribution in China's seas. p. 404-463. Vertebrata. Smithsonian Institution, Florida, USA. 598 p. (Ref. 47843) Pyle, R.0 Chaetodontidae. Butterflyfishes. p.3224-3265. In K.E. Carpenter and V.H. Niem (eds.) FAO species identification guide for fishery purposes. The living marine resources of the Western Central Pacific. Volume 5. Bony fishes part 3 (Menidae to Pomacentridae). Rome, FAO. pp. 2791-3380. (Ref. 48331) Chen, J.-P., K.-T. Shao and C.-P. Lin0 A checklist of reef fishes from the Tungsha Tao (Pratas Island), South China Sea. Acta Zoologica Taiwanica 6(2):13-40. (Ref. 48451) Hardy, J.D. Jr. (comp.)0 Coral reef fish species. NOAANational Oceanographic Data Center. NODC Coral Reef Data and Information Management System. USA. 537 p. (Ref. 48497) Werner, T.B. and G.R. Allen0 A rapid marine biodiversity assessment of the Calamianes Islands, Palawan province, Philippines. RAP Bulletin of Biological Assessment 17. Washington, D.C.:Conservation International. (Ref. 48613) Kuiter, R.H. and T. Tonozuka0 Pictorial guide to Indonesian reef fishes. Part 2. Fusiliers - Dragonets, Caesionidae - Callionymidae. Zoonetics, Australia. 304-622 p. (Ref. 48636) Carl, H.0 Danish fish names. Zoological Museum of Copenhagen. Unpublished. (Ref. 51471) Wantiez, L. and C. Chauvet0 First data on community structure and trophic networks of Uvea coral reef fish assemblages (Wallis and Futuna, South Pacific ocean). Cybium 27(2):83-100. (Ref. 52522) Whiteman, E.A. and I.M. Côté0 Monogamy in marine fishes. Biol. Rev. 79:351-375. (Ref. 52884) Randall, J.E.0 Reef and shore fishes of the South Pacific. New Caledonia to Tahiti and the Pitcairn Islands. University of Hawaii Press, Honolulu, Hawaii. 720 p. (Ref. 54980) Kulbicki, M., N. Guillemot and M. Amand0 A general approach to length-weight relationships for New Caledonian lagoon fishes. Cybium 29(3):235-252. (Ref. 55787) To be filled0 Tubbataha List (Electronic file). Angelique Songco (pers.comm). (Ref. 57038) Chinese Academy of Fishery Sciences0 Chinese aquatic germplasm resources database. http://zzzy.fishinfo.cn (Ref. 58108) Mundy, B.C.0 Checklist of the fishes of the Hawaiian Archipelago. Bishop Mus. Bull. Zool. (6):1-704. (Ref. 58302) Nguyen, N.T. and V.Q. Nguyen0 Biodiversity and living resources of the coral reef fishes in Vietnam marine waters. Science and Technology Publishing House, Hanoi. (Ref. 58534) Hoese, D.F., D.J. Bray, J.R. Paxton and G.R. Allen0 Fishes. In Beasley, O.L. and A. Wells (eds.) Zoological Catalogue of Australia. Volume 35 Australia: ABRS & CSIRO Publishing, 2178 p. (Ref. 75154) Johnson, J.W.0 Annotated checklist of the fishes of Moreton Bay, Queensland, Australia. Memoirs of the Queensland Museum 43(2):709-762. (Ref. 78751) Planes, S., P. Romans and R. Lecomte-Finiger0 Genetic evidence of closed life-cycles for some coral reef fishes within Taiaro Lagoon (Tuamotu Archipelago, French Polynesia). Coral Reefs 17:9-14. (Ref. 79668) Thaman, R.R., T. Fong and A. Balawa0 Ilava Ni Navakavu: Finfishes of Vanua Navakavu, Viti Levu, Fiji Islands. SPRH-FIO Biodiversity and Ethnobiodiversity Report No. 4, The University of the South Pacific, Suva, Fiji Islands. (Ref. 79890) Taquet, M. and A. Diringer0 Poissons de l'Océan Indien et de la mer Rouge. Versailles (France): Éditions Quæ. 527 p. (Ref. 81519) Department of Fisheries Malaysia0 Valid local name of Malaysian marine fishes. Department of Fisheries Malaysia. Ministry of Agriculture and Agro-based Industry. 180 p. (Ref. 85449) Fricke, R., M. Kulbicki and L. Wantiez0 Checklist of the fishes of New Caledonia, and their distribution in the Southwest Pacific Ocean (Pisces). Stuttgarter Beiträge zur Naturkunde A, Neue Serie 4:341-463. (Ref. 86942) Bacchet, P., T. Zysman and Y. Lefèvre0 Guide des poissons de Tahiti et ses îles. Tahiti (Polynésie Francaise): Éditions Au Vent des Îles. 608 p. (Ref. 89972) Allen, G.R. and M.V. Erdmann0 Reef fishes of the East Indies. Perth, Australia: Universitiy of Hawai'i Press, Volumes I-III. Tropical Reef Research. (Ref. 90102) Donaldson, T.J.0 Fishes of the remote Southwest Palau Islands: a zoogeographic perspective. Pac. Sci. 50(3):285-308. (Ref. 99878) Patzner, R.A.0 Reproductive strategies of fish. pp. 311-350. In Rocha, M.J., A. Arukwe and B.G. Kapoor (eds). Fish reproduction: cytology, biology and ecology. Science Publisher, Inc. Oxford. 631 p. (Ref. 101194) Kerr, V.C.0 A rapid marine biodiversity assessment of the coral reefs of the northwest lagoon, between Koumac and Yandé, Province Nord, New Caledonia. BioOne Research Evolved:160-178. (Ref. 101627) Iglésias, T.L., A. Dornburg, M.C. Brandley, M.E. Alfaro and D.L. Warren0 Life in the unthinking depths: energetic constraints on encephalization in marine fishes. J. Evol. Biol. 28(5):1080-1090. (Ref. 104901) Alcala, A.C., J.A. Ingles and A.A. Bucol0 Review of the biodiversity of southern Philippines seas. Philipp. Scient. 45:1-61. (Ref. 106380) Stein, A.0 La pêche côtière aux Australes. p. 215-225 Salvat, B.; Bambridge, T.; Tanret, D.; Petit, J. 2015 Environnement marin des îles Australes, Polynésie française. Institut Récifs Coralliens Pacifique, CRIOBE et The Pew Charitable Trusts Polynésie française. Polynésie française, Tahiti, p. 342. (Ref. 108064) Tsuboi, M., W. van der Bijl, B.T. Kopperud, J. Erritzoe, K.L. Voje, A. Kotrschal, K.E. Yopak, S.P. Collin, A.N. Iwaniuk and N. Kolm0 Breakdown of brain-body allometry and the encephalization of birds and mammals. Nat. Ecol. Evol. 2:1495-1500. (Ref. 118717) Allen, G.R. and M.V. Erdmann0 Coral reef fishes of Timor-Leste. p. 33-82. In G.R. Allen and M.V. Erdmann (eds). A rapid biological assessment of Timor-Leste. Conservation International. (Ref. 118793) Friedlander, A.M., J.E. Caselle, E. Ballesteros, E.K. Brown, A. Turchik and E. Sala0 The real bounty: marine biodiversity in the Pitcairn Islands. PLoS ONE 9(6):e100142. (Ref. 121505) Songco, A.0 Personal communication. Tubbataha species list. Excel file "MyReport_Species_21May2019". (Ref. 121724) University of Western Australia0 Benthic BRUVS data 2019. Video BRUVS from Jessica Meeuwig. (Ref. 122752) Qiu, S., B. Chen, J. Du, K.-H. Loh, J. Liao, X. Liu and W. Yang0 Checklist of the coral fish fauna of Xisha Islands, China. Biodiversity Data Journal 9: e63945. (Ref. 123833) Hazes, B.0 Hazes' underwater photos. Collection of Philippine underwater photos. Unpublished. (Ref. 127519) To, A.W.L., K.S.H. Ching and S.K.H. Shea0 Hong Kong reef fish photo guide. Eco-Education and Resources Centre (Ref. 128523) Coleman, R., J.M. Copus, D.M. Coffey, R.K. Whitton and B.W. Bowen0 Shifting reef fish assemblages along a depth gradient in Pohnpei, Micronesia. PeerJ, 6: e4650. (Ref. 128797) Balisco, R.A.T., V.S. Ticzon, B.R. Samaniego, W.-C. Huang, B.J. Gonzales and T.-Y. Liao0 Marine fishes of Palawan, Philippines: species diversity, new records, and conservation status. Regional Studies in Marine Science 60(2023) 102825:1-10. (Ref. 130287) Price, S.A., S.T. Friedman, K.A. Corn, O. Larouche, K. Brockelsby, A.J. Lee, M. Nagaraj, N.G. Bertrand, M. Danao, M.C. Coyne, J.R. Estrada, R. Friedman, E. Hoeft, M. Iwan, D. Gross, J.H. Kao, B. Landry, M.J. Linares, C. McGlinn and …0 FishShapes v1: Functionally relevant measurements of teleost shape and size on three dimensions. Ecology 103(e3829):1-2. (Ref. 137577) FB-pic-5580-12340 http://purl.org/dc/dcmitype/StillImage image/jpeg John E. Randall 1996-11-18 1999-07-07 http://creativecommons.org/licenses/by-nc/3.0/ This work is licensed under a Creative Commons Attribution-Noncommercial 3.0 Unported License. Image can be freely copied and altered but may be used only for non-commercial purposes. Original author and source must be properly acknowledged WorldFish Center - FishBase General public http://www.fishbase.org/summary/SpeciesSummary.php?id=5580 http://rs.tdwg.org/ontology/voc/SPMInfoItems#Distribution http://www.fishbase.us/images/species/Chuli_u1.jpg http://www.fishbase.us/images/thumbnails/jpg/tn_Chuli_u1.jpg FB-pic-5580-12341 http://purl.org/dc/dcmitype/StillImage image/jpeg John E. Randall 1996-11-18 1999-07-09 http://creativecommons.org/licenses/by-nc/3.0/ This work is licensed under a Creative Commons Attribution-Noncommercial 3.0 Unported License. Image can be freely copied and altered but may be used only for non-commercial purposes. Original author and source must be properly acknowledged http://www.fishbase.org/summary/SpeciesSummary.php?id=5580 http://www.fishbase.us/images/species/Chuli_u2.jpg http://www.fishbase.us/images/thumbnails/jpg/tn_Chuli_u2.jpg FB-GeneralDescr-5580 http://purl.org/dc/dcmitype/Text text/html Estelita Emily Capuli 1992-06-05 00:00:00 2025-11-20 00:00:00 Biology http://creativecommons.org/licenses/by-nc/3.0/ WorldFish Center - FishBase General public http://www.fishbase.org/summary/SpeciesSummary.php?id=5580 http://rs.tdwg.org/ontology/voc/SPMInfoItems#LifeCycle Adults occur in coral-rich areas of lagoon reefs and less commonly in seaward reefs. Juveniles in harbors and estuaries (Ref. 48636). Benthopelagic (Ref. 58302). Usually solitary, in pairs or in small groups. Adults feed on plant and animal material. Oviparous (Ref. 205), monogamous (Ref. 52884). Form pairs during breeding (Ref. 205). Can be easily maintained in tanks. Myers, R.F. 1991 Micronesian reef fishes. Second Ed. Coral Graphics, Barrigada, Guam. 298 p. (Ref. 1602) FB-DiagnosticDescr-5580 http://purl.org/dc/dcmitype/Text text/html Rodolfo B. Reyes 1994-01-13 00:00:00 2013-04-18 00:00:00 http://creativecommons.org/licenses/by-nc/3.0/ WorldFish Center - FishBase General public http://www.fishbase.org/summary/SpeciesSummary.php?id=5580 http://rs.tdwg.org/ontology/voc/SPMInfoItems#Size Description: Easily recognized on color and distinguished from sibling by white color between the black saddles (Ref. 48636). Snout length 2.2-2.5 in HL. Body depth 1.5-1.8 in SL (Ref. 90102). Burgess, W.E. 1978 Butterflyfishes of the world. A monograph of the Family Chaetodontidae. T.F.H. Publications, Neptune City, New Jersey. (Ref. 4855) FB-Morphology-5580 http://purl.org/dc/dcmitype/Text text/html Rodolfo B. Reyes 1994-01-13 00:00:00 2013-04-18 00:00:00 http://creativecommons.org/licenses/by-nc/3.0/ WorldFish Center - FishBase General public http://www.fishbase.org/summary/SpeciesSummary.php?id=5580 http://rs.tdwg.org/ontology/voc/SPMInfoItems#Threats Dorsal spines (total): 12; Dorsal soft rays (total): 23 - 25; Anal spines: 3 - 4; Analsoft rays: 19 - 21 Burgess, W.E. 1978 Butterflyfishes of the world. A monograph of the Family Chaetodontidae. T.F.H. Publications, Neptune City, New Jersey. (Ref. 4855) FB-Distribution-5580 http://purl.org/dc/dcmitype/Text text/html Estelita Emily Capuli 1992-06-05 00:00:00 2025-09-05 00:00:00 http://creativecommons.org/licenses/by-nc/3.0/ WorldFish Center - FishBase General public http://www.fishbase.org/summary/SpeciesSummary.php?id=5580 http://rs.tdwg.org/ontology/voc/SPMInfoItems#Habitat Indo-Pacific: Cocos-Keeling Islands to the Tuamoto Islands, north to Japan. Randall, J.E., G.R. Allen and R.C. Steene 1990 Fishes of the Great Barrier Reef and Coral Sea. University of Hawaii Press, Honolulu, Hawaii. 506 p. (Ref. 2334) FB-Uses-5580 http://purl.org/dc/dcmitype/Text text/html Estelita Emily Capuli 1992-06-05 00:00:00 2025-11-20 00:00:00 Importance http://creativecommons.org/licenses/by-nc/3.0/ WorldFish Center - FishBase General public http://www.fishbase.org/summary/SpeciesSummary.php?id=5580 http://rs.tdwg.org/ontology/voc/SPMInfoItems#Distribution fisheries: minor commercial; aquarium: commercial; price category: unknown; price reliability: Edwards, A.J. and A.D. Shepherd 1992 Environmental implications of aquarium-fish collection in the Maldives, with proposals for regulation. Environ. Conserv. 19:61-72. (Ref. 4907) FB-LifeCycle-5580 http://purl.org/dc/dcmitype/Text text/html Susan M. Luna 1992-06-05 00:00:00 2025-11-20 00:00:00 http://creativecommons.org/licenses/by-nc/3.0/ http://www.fishbase.org//Reproduction/FishReproSummary.php?ID=5580&StockCode=5870 Distinct pairing (Ref. 205). Monogamous mating is observed as both obligate and social (Ref. 52884). Thresher, R.E. 1984 Reproduction in reef fishes. T.F.H. Publications, Inc. Ltd., Neptune City, New Jersey. 399 p. (Ref. 240) FB-TrophicStrategy-5580 http://purl.org/dc/dcmitype/Text text/html Susan M. Luna 1992-12-01 00:00:00 2008-12-08 00:00:00 http://creativecommons.org/licenses/by-nc/3.0/ WorldFish Center - FishBase General public ulietensis http://rs.tdwg.org/ontology/voc/SPMInfoItems#LifeCycle Occurs in coral-rich areas of lagoon reefs and less commonly in seaward reefs. Usually solitary, in pairs or in small groups. Feeds on plant and animal material. Can be easily maintained in tanks. Steene, R.C. 1978 Butterfly and angelfishes of the world. A.H. & A.W. Reed Pty Ltd., Australia. vol. 1. 144 p. (Ref. 4859) FB-Size-5580 http://purl.org/dc/dcmitype/Text text/html Estelita Emily Capuli 1992-06-05 00:00:00 2025-11-20 00:00:00 Max. size http://creativecommons.org/licenses/by-nc/3.0/ WorldFish Center - FishBase General public http://www.fishbase.org/summary/SpeciesSummary.php?id=5580 http://rs.tdwg.org/ontology/voc/SPMInfoItems#Size 15.0 cm TL (male/unsexed; (Ref. 4859)) Steene, R.C. 1978 Butterfly and angelfishes of the world. A.H. & A.W. Reed Pty Ltd., Australia. vol. 1. 144 p. (Ref. 4859) FB-Threats-2 http://purl.org/dc/dcmitype/Text text/html Belen Acosta 1990-10-18 00:00:00 2024-03-07 00:00:00 http://creativecommons.org/licenses/by-nc/3.0/ WorldFish Center - FishBase General public http://www.fishbase.org/summary/SpeciesSummary.php?id=2 http://rs.tdwg.org/ontology/voc/SPMInfoItems#Threats Least Concern (LC) , IUCN Grouper and Wrasse Specialist Group IUCN 2006 2006 IUCN red list of threatened species. www.iucnredlist.org. Downloaded July 2006. FB-Habitat-5580 http://purl.org/dc/dcmitype/Text text/html Estelita Emily Capuli 1992-06-05 00:00:00 2025-11-20 00:00:00 Environment http://creativecommons.org/licenses/by-nc/3.0/ WorldFish Center - FishBase General public http://www.fishbase.org/summary/SpeciesSummary.php?id=5580 http://rs.tdwg.org/ontology/voc/SPMInfoItems#Habitat reef-associated; non-migratory (Ref. 52884); marine; depth range 0 - 40 m (Ref. 128797) Whiteman, E.A. and I.M. Côté 2004 Monogamy in marine fishes. Biol. Rev. 79:351-375. (Ref. 52884) Coleman, R., J.M. Copus, D.M. Coffey, R.K. Whitton and B.W. Bowen 2018 Shifting reef fish assemblages along a depth gradient in Pohnpei, Micronesia. PeerJ, 6: e4650. (Ref. 128797)