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	<dc:identifier>FB-5508</dc:identifier>
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	<dwc:Kingdom>Animalia</dwc:Kingdom>
	<dwc:Phylum>Chordata</dwc:Phylum>
	<dwc:Class>Elasmobranchii</dwc:Class>
	<dwc:Order>Myliobatiformes</dwc:Order>
	<dwc:Family>Gymnuridae</dwc:Family>
	<dwc:Genus>Gymnura</dwc:Genus>
	<dwc:ScientificName>Gymnura natalensis (Gilchrist &amp; Thompson, 1911)</dwc:ScientificName>
	<commonName xml:lang='af'>Rem-vinderrog</commonName>
	<commonName xml:lang='af'>Rem-vlinderrog</commonName>
	<commonName xml:lang='de'>Schmetterlingsrochen</commonName>
	<commonName xml:lang='en'>Backwater butterfly ray</commonName>
	<commonName xml:lang='en'>Backwater butterflyray</commonName>
	<commonName xml:lang='en'>Butterfly ray</commonName>
	<commonName xml:lang='en'>Diamond ray</commonName>
	<commonName xml:lang='et'>Natali liblikrai</commonName>
	<commonName xml:lang='fr'>Raie-papillon du Natal</commonName>
	<commonName xml:lang='es'>Raya mariposa de Natal</commonName>
	<commonName xml:lang=''>Diamantvlinderrog</commonName>
	<commonName xml:lang=''>南非沙粒魟</commonName>
	<commonName xml:lang=''>南非燕魟</commonName>
	<synonym relationship='synonym'>Pteroplatea natalensis Gilchrist &amp; Thompson, 1911</synonym>
	<synonym relationship='synonym'>Urogymnus natalensis (Gilchrist &amp; Thompson, 1911)</synonym>
	<dcterms:created>1992-06-02</dcterms:created>
	<dcterms:modified>2024-08-16</dcterms:modified>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=205&amp;speccode=5508'>Breder, C.M. and D.E. Rosen0 Modes of reproduction in fishes. T.F.H. Publications, Neptune City, New Jersey. 941 p. (Ref. 205)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=557&amp;speccode=5508'>Wheeler, A.0 Das grosse Buch der Fische. Eugen Ulmer GmbH &amp; Co. Stuttgart. 356 p. (Ref. 557)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=3263&amp;speccode=5508'>Compagno, L.J.V.0 Dasyatidae. p. 135-142. In M.M. Smith and P.C. Heemstra (eds.) Smiths' sea fishes. Springer-Verlag, Berlin. (Ref. 3263)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=3670&amp;speccode=5508'>van der Elst, R.0 A guide to the common sea fishes of southern Africa. C. Struik, Cape Town. 367 p. (Ref. 3670)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=3678&amp;speccode=5508'>Torres, F.S.B. Jr.0 Tabular data on marine fishes from Southern Africa, Part I. Length-weight relationships. Fishbyte 9(1):50-53. (Ref. 3678)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=4332&amp;speccode=5508'>van der Elst, R.P. and F. Adkin (eds.)0 Marine linefish: priority species and research objectives in southern Africa. Oceanogr. Res. Inst., Spec. Publ. No.1. 132 p. (Ref. 4332)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=5285&amp;speccode=5508'>Smith, J.L.B.0 Fishes of Inhaca. p. 131-136. In W. Macnae and M. Kalk (eds.) A natural history of Inhaca Island, Moçambique. Witwatersrand University Press, Johannesburg. (Ref. 5285)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=5578&amp;speccode=5508'>Compagno, L.J.V., D.A. Ebert and M.J. Smale0 Guide to the sharks and rays of southern Africa. New Holland (Publ.) Ltd., London. 158 p. (Ref. 5578)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=11228&amp;speccode=5508'>Heemstra, P.C.0 Additions and corrections for the 1995 impression. p. v-xv. In M.M. Smith and P.C. Heemstra (eds.) Revised Edition of Smiths' Sea Fishes. Springer-Verlag, Berlin. (Ref. 11228)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=12484&amp;speccode=5508'>van der Elst, R.0 A guide to the common sea fishes of southern Africa. (3rd Ed.). Struik Publishers, Cape Town. 398 p. (Ref. 12484)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=27121&amp;speccode=5508'>Bianchi, G., K.E. Carpenter, J.-P. Roux, F.J. Molloy, D. Boyer and H.J. Boyer0 FAO species identification guide for fishery purposes. Field guide to the living marine resources of Namibia. FAO, Rome. 265p. (Ref. 27121)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=28016&amp;speccode=5508'>al Sakaff, H. and M. Esseen0 Occurrence and distribution of fish species off Yemen (Gulf of Aden and Arabian Sea). Naga, ICLARM Q. 22(1):43-47. (Ref. 28016)</reference>
	<reference isbn='957-8596-59-6' url='http://www.fishbase.org/references/FBRefSummary.php?id=31517&amp;speccode=5508'>Wu, H.L., K.-T. Shao and C.F. Lai (eds.)0 Latin-Chinese dictionary of fishes names. The Sueichan Press, Taiwan. 1028 p. (Ref. 31517)</reference>
	<reference isbn='0-8018-6048-2' url='http://www.fishbase.org/references/FBRefSummary.php?id=35766&amp;speccode=5508'>Compagno, L.J.V.0 Checklist of living elasmobranchs. p. 471-498. In W.C. Hamlett (ed.) Sharks, skates, and rays: the biology of elasmobranch fishes. Johns Hopkins University Press, Maryland. (Ref. 35766)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=36670&amp;speccode=5508'>Anonymous0 Fish collection database of the J.L.B. Smith Institute of Ichthyology, Grahamstown, South Africa. J.L.B. Smith Institute of Ichthyology, Grahamstown, South Africa. (Ref. 36670)</reference>
	<reference isbn='92-5-104345-0' url='http://www.fishbase.org/references/FBRefSummary.php?id=36731&amp;speccode=5508'>Bianchi, G., K.E. Carpenter, J.-P. Roux, F.J. Molloy, D. Boyer and H.J. Boyer0 FAO species identification guide for fishery purposes. Field guide to the living marine resources of Namibia. FAO, Rome. 265 p. (Ref. 36731)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=40637&amp;speccode=5508'>IGFA0 Database of IGFA angling records until 2001. IGFA, Fort Lauderdale, USA. (Ref. 40637)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=43454&amp;speccode=5508'>Bor, P.H.F.0 Nederlandse naamlijst van de recente haaien en roggen (Chondrichthyes: Elasmobranchii) van de wereld. World Wide Web electronic publication www.rajidae.tmfweb.nl, version (05/2002). (Ref. 43454)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=50449&amp;speccode=5508'>Dulvy, N.K. and J.D. Reynolds0 Evolutionary transitions among egg-laying, live-bearing and maternal inputs in sharks and rays. Proc. R. Soc. Lond., Ser. B: Biol. Sci. 264:1309-1315. (Ref. 50449)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=58108&amp;speccode=5508'>Chinese Academy of Fishery Sciences0 Chinese aquatic germplasm resources database. http://zzzy.fishinfo.cn (Ref. 58108)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=75549&amp;speccode=5508'>FAO-FIES0 Aquatic Sciences and Fisheries Information System (ASFIS) species list. Retrieved from http://www.fao.org/fishery/collection/asfis, 29 April 2008. (Ref. 75549)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=83882&amp;speccode=5508'>FAO-FIES0 Aquatic Sciences and Fisheries Information System (ASFIS) species list. Retrieved from http://www.fao.org/fishery/collection/asfis/en, March 2010. (Ref. 83882)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=90062&amp;speccode=5508'>FAO-FIES0 Aquatic Sciences and Fisheries Information System (ASFIS) species list. Retrieved from http://www.fao.org/fishery/collection/asfis/en, March 2012. (Ref. 90062)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=95632&amp;speccode=5508'>FAO-FIES0 Aquatic Sciences and Fisheries Information System (ASFIS) species list. Retrieved from http://www.fao.org/fishery/collection/asfis/en, April 2014. (Ref. 95632)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=101110&amp;speccode=5508'>FAO-FIES0 Aquatic Sciences and Fisheries Information System (ASFIS) species list. Retrieved from http://www.fao.org/fishery/collection/asfis/en, [accessed 13/04/2015]. (Ref. 101110)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=105471&amp;speccode=5508'>eElurikkus0 eElurikkus koondab ühte portaali Eesti eluslooduse andmed. http://elurikkus.ut.ee/ (Ref. 105471)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=115257&amp;speccode=5508'>FAO-FIES0 Aquatic Sciences and Fisheries Information System (ASFIS) species list. Retrieved from http://www.fao.org/fishery/collection/asfis/en (accessed 08/06/2017). (Ref. 115257)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=118272&amp;speccode=5508'>FAO-FIES0 Aquatic Sciences and Fisheries Information System (ASFIS) species list. Retrieved from htttps://www.fao.org/fishery/collection/asfis/en [accessed May, 2018] (Ref. 118272)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=129684&amp;speccode=5508'>Daly, R., G. Jordaan, D. Parker, G. Cliff, N. Nkabi, R. Kyle, S. Fennessy and B. Mann0 Movement patterns and catch trends of the diamond ray &lt;i&gt;Gymnura natalensis&lt;/i&gt; (Dasyatidae) in South African waters. African Journal of Marine Science 44(1):35-48. (Ref. 129684)</reference><dataObject>
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	<agent homepage='' logoURL='' role='recorder'>Crispina B. Binohlan</agent>
	<dcterms:created>1992-06-02 00:00:00</dcterms:created>
	<dcterms:modified>2024-08-16 00:00:00</dcterms:modified>
	<dc:title>Biology</dc:title>
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	<dc:description>Occurs off sandy beaches, muddy estuaries and offshore banks.  Found singly or in large groups (Ref. 5578).  Feeds on a variety of fishes, crabs and polychaete worms (Ref. 5578).  Ovoviviparous (Ref. 50449).  Ability to change markings and color to blend into environment (Ref. 3263).  Caught by offshore trawlers (Ref. 5578).  Caught by shore anglers, it is prized for its strong fight when hooked, often released (Ref. 5578).</dc:description>
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	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=3263&amp;speccode=5508'>Compagno, L.J.V. 1986 Dasyatidae. p. 135-142. In M.M. Smith and P.C. Heemstra (eds.) Smiths' sea fishes. Springer-Verlag, Berlin. (Ref. 3263)</reference></dataObject><dataObject>
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	<agent homepage='' logoURL='' role='recorder'>Cristina V. Garilao</agent>
	<dcterms:created>1996-11-28 00:00:00</dcterms:created>
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	<dc:description>Broad diamond-shaped pectoral disc almost twice as wide as it is long; tail shorter than body with black and white bands and a small sting; tentacle at rear edge of each spiracle (Ref. 5578).  Grey, green or brown above, often with darker mottling; white below; can change color of upper disc rapidly to match substrate (Ref. 5578).</dc:description>
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	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=5578&amp;speccode=5508'>Compagno, L.J.V., D.A. Ebert and M.J. Smale 1989 Guide to the sharks and rays of southern Africa. New Holland (Publ.) Ltd., London. 158 p. (Ref. 5578)</reference></dataObject><dataObject>
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	<dcterms:created>1996-11-28 00:00:00</dcterms:created>
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	<dc:description>Dorsal spines (total): 0; Dorsal soft rays (total): 0</dc:description>
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	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=5578&amp;speccode=5508'>Compagno, L.J.V., D.A. Ebert and M.J. Smale 1989 Guide to the sharks and rays of southern Africa. New Holland (Publ.) Ltd., London. 158 p. (Ref. 5578)</reference></dataObject><dataObject>
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	<agent homepage='' logoURL='' role='recorder'>Crispina B. Binohlan</agent>
	<dcterms:created>1992-06-02 00:00:00</dcterms:created>
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	<subject>http://rs.tdwg.org/ontology/voc/SPMInfoItems#Size</subject>
	<dc:description>Southeast Atlantic and Western Indian Ocean:  Namibia round the Cape to southern Mozambique.</dc:description>
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	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=5578&amp;speccode=5508'>Compagno, L.J.V., D.A. Ebert and M.J. Smale 1989 Guide to the sharks and rays of southern Africa. New Holland (Publ.) Ltd., London. 158 p. (Ref. 5578)</reference></dataObject><dataObject>
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	<dc:title>Importance</dc:title>
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	<subject>http://rs.tdwg.org/ontology/voc/SPMInfoItems#Threats</subject>
	<dc:description>fisheries: bycatch; gamefish: yes</dc:description>
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	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=4332&amp;speccode=5508'>van der Elst, R.P. and F. Adkin (eds.) 1991 Marine linefish: priority species and research objectives in southern Africa. Oceanogr. Res. Inst., Spec. Publ. No.1. 132 p. (Ref. 4332)</reference></dataObject><dataObject>
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	<dc:description>Exhibit ovoviparity (aplacental viviparity), with embryos feeding initially on yolk, then receiving additional nourishment from the mother by indirect absorption of uterine fluid enriched with mucus, fat or protein through specialised structures (Ref. 50449). Distinct pairing with embrace (Ref. 205).  One year gestation; up to 10 young (Ref. 5578).</dc:description>
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	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=205&amp;speccode=5508'>Breder, C.M. and D.E. Rosen 1966 Modes of reproduction in fishes. T.F.H. Publications, Neptune City, New Jersey. 941 p. (Ref. 205)</reference></dataObject><dataObject>
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	<agent homepage='' logoURL='' role='recorder'>Pascualita Sa-a</agent>
	<dcterms:created>1999-09-22 00:00:00</dcterms:created>
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	<dc:source>natalensis</dc:source>
	<subject>http://rs.tdwg.org/ontology/voc/SPMInfoItems#Distribution</subject>
	<dc:description>Feeds on fish and zoobenthos (Ref. 5578).</dc:description>
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	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=5578&amp;speccode=5508'>Compagno, L.J.V., D.A. Ebert and M.J. Smale 1989 Guide to the sharks and rays of southern Africa. New Holland (Publ.) Ltd., London. 158 p. (Ref. 5578)</reference></dataObject><dataObject>
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	<dc:description>250 cm WD (male/unsexed; (Ref. 3263)); max. published weight: 82.6 kg (Ref. 40637)</dc:description>
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	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=3263&amp;speccode=5508'>Compagno, L.J.V. 1986 Dasyatidae. p. 135-142. In M.M. Smith and P.C. Heemstra (eds.) Smiths' sea fishes. Springer-Verlag, Berlin. (Ref. 3263)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=40637&amp;speccode=5508'>IGFA 2001 Database of IGFA angling records until 2001. IGFA, Fort Lauderdale, USA. (Ref. 40637)</reference></dataObject><dataObject>
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	<dc:rights></dc:rights>
	<dcterms:rightsHolder>WorldFish Center - FishBase</dcterms:rightsHolder>
	<dcterms:bibliographicCitation></dcterms:bibliographicCitation>
	<audience>General public</audience>
	<dc:source>http://www.fishbase.org/summary/SpeciesSummary.php?id=2</dc:source>
	<subject>http://rs.tdwg.org/ontology/voc/SPMInfoItems#Size</subject>
	<dc:description>Least Concern (LC) , IUCN Grouper and Wrasse Specialist Group</dc:description>
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	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=57073'>IUCN 2006 2006 IUCN red list of threatened species. www.iucnredlist.org. Downloaded July 2006.</reference></dataObject><dataObject>
	<dc:identifier>FB-Habitat-5508</dc:identifier>
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	<agent homepage='' logoURL='' role='recorder'>Crispina B. Binohlan</agent>
	<dcterms:created>1992-06-02 00:00:00</dcterms:created>
	<dcterms:modified>2024-08-16 00:00:00</dcterms:modified>
	<dc:title>Environment</dc:title>
	<dc:language></dc:language>
	<license>http://creativecommons.org/licenses/by-nc/3.0/</license>
	<dc:rights></dc:rights>
	<dcterms:rightsHolder>WorldFish Center - FishBase</dcterms:rightsHolder>
	<dcterms:bibliographicCitation></dcterms:bibliographicCitation>
	<audience>General public</audience>
	<dc:source>http://www.fishbase.org/summary/SpeciesSummary.php?id=5508</dc:source>
	<subject>http://rs.tdwg.org/ontology/voc/SPMInfoItems#Threats</subject>
	<dc:description>demersal; marine; depth range 20 - 100 m (Ref. 28016)</dc:description>
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	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=28016&amp;speccode=5508'>al Sakaff, H. and M. Esseen 1999 Occurrence and distribution of fish species off Yemen (Gulf of Aden and Arabian Sea). Naga, ICLARM Q. 22(1):43-47. (Ref. 28016)</reference></dataObject>
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