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	<dc:identifier>FB-5416</dc:identifier>
	<dc:source>http://www.fishbase.org/summary/SpeciesSummary.php?id=5416</dc:source>
	<dwc:Kingdom>Animalia</dwc:Kingdom>
	<dwc:Phylum>Chordata</dwc:Phylum>
	<dwc:Class>Teleostei</dwc:Class>
	<dwc:Order>Cypriniformes</dwc:Order>
	<dwc:Family>Cyprinidae</dwc:Family>
	<dwc:Genus>Labeobarbus</dwc:Genus>
	<dwc:ScientificName>Labeobarbus johnstonii (Boulenger, 1907)</dwc:ScientificName>
	<commonName xml:lang='cs'>Parma Johnstonova</commonName>
	<commonName xml:lang='cs'>Parma široústá</commonName>
	<commonName xml:lang='et'>Johnstoni mokkparrak</commonName>
	<commonName xml:lang='et'>Laisuu-pardkala</commonName>
	<commonName xml:lang='et'>Lühisaba-mokkparrak</commonName>
	<commonName xml:lang='sw'>Chikasu</commonName>
	<commonName xml:lang='sw'>Kadyakolo</commonName>
	<commonName xml:lang='sw'>Kuju</commonName>
	<commonName xml:lang='sw'>Ngumbo</commonName>
	<commonName xml:lang='sw'>Nkumbwa</commonName>
	<commonName xml:lang=''>Mpondo</commonName>
	<commonName xml:lang=''>Ngumbo</commonName>
	<commonName xml:lang=''>Kadyakolo</commonName>
	<commonName xml:lang=''>Matemba</commonName>
	<commonName xml:lang=''>約氏魮</commonName>
	<commonName xml:lang=''>约氏魮</commonName>
	<commonName xml:lang=''>闊嘴魮</commonName>
	<commonName xml:lang=''>阔嘴魮</commonName>
	<synonym relationship='synonym'>Barbus eurystomus Keilhack, 1908</synonym>
	<synonym relationship='synonym'>Barbus globiceps Worthington, 1933</synonym>
	<synonym relationship='synonym'>Barbus intermedius brevicauda Keilhack, 1908</synonym>
	<synonym relationship='synonym'>Barbus intermedius eurystomus Keilhack, 1908</synonym>
	<synonym relationship='synonym'>Barbus johnstonii Boulenger, 1907</synonym>
	<synonym relationship='synonym'>Barbus njassae Keilhack, 1908</synonym>
	<synonym relationship='synonym'>Barbus nyassae Keilhack, 1908</synonym>
	<synonym relationship='synonym'>Labeobarbus brevicauda (Keilhack, 1908)</synonym>
	<synonym relationship='synonym'>Varicorhinus nyasensis Worthington, 1933</synonym>
	<dcterms:created>1992-04-20</dcterms:created>
	<dcterms:modified>2024-01-25</dcterms:modified>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=2801&amp;speccode=5416'>Lévêque, C. and J. Daget0 Cyprinidae. p. 217-342. In J. Daget, J.-P. Gosse and D.F.E. Thys van den Audenaerde (eds.) Check-list of the freshwater fishes of Africa (CLOFFA). ORSTOM, Paris and MRAC, Tervuren. Vol. 1. (Ref. 2801)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=4364&amp;speccode=5416'>Jackson, P.B.N.0 Check list of the fishes of Nyasaland. The National Museum of Southern Rhodesia. Occasional Papers 3(25B). (Ref. 4364)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=4713&amp;speccode=5416'>Bernacsek, G.M., A. Massinga and P. Contreras0 Mozambique. Exploratory gill-netting in Lake Niassa, Mozambique, with biological profile of the main taxa caught. A report prepared for the Development of Inland and Inshore Fisheries (Phase II) Project. FAO/GDP/MOZ/006/SWE (Phase II) Field Doc. 5. (Ref. 4713)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=4967&amp;speccode=5416'>Eccles, D.H.0 FAO species identification sheets for fishery purposes. Field guide to the freshwater fishes of Tanzania. Prepared and published with the support of the United Nations Development Programme (project URT/87/016). FAO, Rome. 145 p. (Ref. 4967)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=5595&amp;speccode=5416'>Konings, A.0 Ad Konings's book of cichlids and all the other fishes of Lake Malawi. T.F.H. Publications, Inc. 495 p. (Ref. 5595)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=12818&amp;speccode=5416'>Anonymous0 Fish registrations within the museum database of the Vertebrate Section of the Royal Museum for Central Africa. MRAC, Tervuren, Belgium. (Ref. 12818)</reference>
	<reference isbn='957-8596-59-6' url='http://www.fishbase.org/references/FBRefSummary.php?id=31517&amp;speccode=5416'>Wu, H.L., K.-T. Shao and C.F. Lai (eds.)0 Latin-Chinese dictionary of fishes names. The Sueichan Press, Taiwan. 1028 p. (Ref. 31517)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=31982&amp;speccode=5416'>Anonymous0 Fish collection database of the Natural History Museum, London (formerly British Museum of Natural History (BMNH)). Natural History Museum, London (formerly British Museum of Natural History (BMNH)). (Ref. 31982)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=33021&amp;speccode=5416'>Eschmeyer, W.N. (ed.)0 Catalog of fishes. Updated database version of November 1999. Catalog databases as made available to FishBase in November 1999. (Ref. 33021)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=33486&amp;speccode=5416'>Nsiku, E.0 Changes in the fisheries of Lake Malawi, 1976-1996: ecosystem-based analysis. University of British Columbia, Vancouver, Canada. 217 p. M.Sc. thesis. (Ref. 33486)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=36653&amp;speccode=5416'>Anonymous0 Fish collection database of the National Museum of Zimbabwe. National Museum of Zimbabwe, P.O. Box 240, Bulawayo, Zimbabwe. (Ref. 36653)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=36670&amp;speccode=5416'>Anonymous0 Fish collection database of the J.L.B. Smith Institute of Ichthyology, Grahamstown, South Africa. J.L.B. Smith Institute of Ichthyology, Grahamstown, South Africa. (Ref. 36670)</reference>
	<reference isbn='0-86810-003-X' url='http://www.fishbase.org/references/FBRefSummary.php?id=37112&amp;speccode=5416'>Tweddle, D., D.S.C. Lewis and N.G. Willoughby0 The nature of the barrier separating the Lake Malawi and Zambezi fish faunas. Ichthyol. Bull. Rhodes Univ. 39:1-9. (Ref. 37112)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=38732&amp;speccode=5416'>Anonymous0 Fish collection database of the National Museum of Natural History (Smithsonian Institution). Smithsonian Institution-Division of Fishes. (Ref. 38732)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=41414&amp;speccode=5416'>Anonymous0 Fish collection database of the American Museum of Natural History. American Museum of Natural History, Central Park West , NY 10024-5192, USA. (Ref. 41414)</reference>
	<reference isbn='80-7036-139-5' url='http://www.fishbase.org/references/FBRefSummary.php?id=51055&amp;speccode=5416'>Hanel, L. and J. Novák0 České názvy zivočichů V. Ryby a ryboviti obratlovci (Pisces) 3., maloústí (Gonorhynchiformes) - máloostní (Cypriniformes). Národní muzeum (zoologické oddělení), Praha. (Ref. 51055)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=51243&amp;speccode=5416'>Riede, K.0 Global register of migratory species - from global to regional scales. Final Report of the R&amp;D-Projekt 808 05 081. Federal Agency for Nature Conservation, Bonn, Germany. 329 p. (Ref. 51243)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=54621&amp;speccode=5416'>Eschmeyer, W.N. (ed.)0 Catalog of fishes. Updated database version of May 2005. Database dump. Catalog databases as made available to FishBase in May 2005. (Ref. 54621)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=55463&amp;speccode=5416'>Snoeks, J. (ed.)0 The cichlid diversity of Lake Malawi/Nyasa/Niassa: identification, distribution and taxonomy. Cichlid Press, El Paso, USA, 360p. (Ref. 55463)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=55909&amp;speccode=5416'>Snoeks, J.0 The non-cichlid fishes of the Lake Malawi system: a compilation. p. 20-26. In Snoeks, J. (ed.) The cichlid diversity of Lake Malawi/Nyasa/Niassa: identification, distribution and taxonomy. Cichlid Press, El Paso, USA, 360p. (Ref. 55909)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=58108&amp;speccode=5416'>Chinese Academy of Fishery Sciences0 Chinese aquatic germplasm resources database. http://zzzy.fishinfo.cn (Ref. 58108)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=75045&amp;speccode=5416'>Tweddle, D. and P.H. Skelton0 New species of &lt;i&gt;Barbus&lt;/i&gt; and &lt;i&gt;Labeobarbus&lt;/i&gt; (Teleostei: Cyprinidae) from the South Rukuru River, Malawi, Africa. Smithiana Bulletin 8:25-39. (Ref. 75045)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=83882&amp;speccode=5416'>FAO-FIES0 Aquatic Sciences and Fisheries Information System (ASFIS) species list. Retrieved from http://www.fao.org/fishery/collection/asfis/en, March 2010. (Ref. 83882)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=90062&amp;speccode=5416'>FAO-FIES0 Aquatic Sciences and Fisheries Information System (ASFIS) species list. Retrieved from http://www.fao.org/fishery/collection/asfis/en, March 2012. (Ref. 90062)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=95632&amp;speccode=5416'>FAO-FIES0 Aquatic Sciences and Fisheries Information System (ASFIS) species list. Retrieved from http://www.fao.org/fishery/collection/asfis/en, April 2014. (Ref. 95632)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=101110&amp;speccode=5416'>FAO-FIES0 Aquatic Sciences and Fisheries Information System (ASFIS) species list. Retrieved from http://www.fao.org/fishery/collection/asfis/en, [accessed 13/04/2015]. (Ref. 101110)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=105471&amp;speccode=5416'>eElurikkus0 eElurikkus koondab ühte portaali Eesti eluslooduse andmed. http://elurikkus.ut.ee/ (Ref. 105471)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=108777&amp;speccode=5416'>Vreven, E.J.W.M.N., T. Musschoot, J. Snoeks and U.K. Schliewen0 The African hexaploid Torini (Cypriniformes: Cyprinidae): review of a tumultuous history. Zool. J. Lin. Soc. 177:231-305. (Ref. 108777)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=114499&amp;speccode=5416'>Boulenger, G.A.0 Descriptions of three new fishes from Central Africa. Ann. Mag. Nat. Hist. (Ser. 7) 20:487-489. (Ref. 114499)</reference>
	<reference isbn='1849954984' url='http://www.fishbase.org/references/FBRefSummary.php?id=128868&amp;speccode=5416'>Beolens, B., M. Grayson and M. Watkins0 Eponym dictionary of Fishes. 	Dunbeath, Caithness (Scotland, UK): Whittles Publishing Limited, 1300 p. (Ref. 128868)</reference><dataObject>
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	<agent homepage='' logoURL='' role='recorder'>Susan M. Luna</agent>
	<dcterms:created>1992-04-20 00:00:00</dcterms:created>
	<dcterms:modified>2024-01-25 00:00:00</dcterms:modified>
	<dc:title>Biology</dc:title>
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	<dc:description>Inhabits inshore areas of the lake, but ascends rivers to breed (Ref. 4967). Found in all types of habitat and occurs in groups of about 25 individuals; not exported for trade (Ref. 5595).</dc:description>
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	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=2801&amp;speccode=5416'>Lévêque, C. and J. Daget 1984 Cyprinidae. p. 217-342. In J. Daget, J.-P. Gosse and D.F.E. Thys van den Audenaerde (eds.) Check-list of the freshwater fishes of Africa (CLOFFA). ORSTOM, Paris and MRAC, Tervuren. Vol. 1. (Ref. 2801)</reference></dataObject><dataObject>
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	<dc:description>Diagnosis: High lateral line scale count, 31-40 scales on lateral line; long, robust barbelsof equal length; and enlarged, molariform pharyngeal teeth (Ref. 75045).Description: Snout rounded; complete range of variation between the soft-lipped form and the shovel-lipped form (Ref. 4967). A high gillraker count, 10-15 gill rakers (Ref. 75045). Dorsal fin with 4-5 unbranched rays; a flexible and not heavily ossified last simple dorsal ray (Ref. 75045). Predorsal scale rows 10-14 (Ref. 75045). Total vertebrae 40-43; 22-24 abdominal vertebrae, 16-19 caudal vertebrae and 10-12 predorsal vertebrae; 14-16 rib pairs (Ref. 75045).</dc:description>
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	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=4967&amp;speccode=5416'>Eccles, D.H. 1992 FAO species identification sheets for fishery purposes. Field guide to the freshwater fishes of Tanzania. Prepared and published with the support of the United Nations Development Programme (project URT/87/016). FAO, Rome. 145 p. (Ref. 4967)</reference></dataObject><dataObject>
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	<dc:description>Dorsal spines (total): 0; Dorsal soft rays (total): 12 - 14;  Analspines: 0; Analsoft rays: 8 - 9; Vertebrae: 40 - 43</dc:description>
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	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=4967&amp;speccode=5416'>Eccles, D.H. 1992 FAO species identification sheets for fishery purposes. Field guide to the freshwater fishes of Tanzania. Prepared and published with the support of the United Nations Development Programme (project URT/87/016). FAO, Rome. 145 p. (Ref. 4967)</reference></dataObject><dataObject>
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	<subject>http://rs.tdwg.org/ontology/voc/SPMInfoItems#Habitat</subject>
	<dc:description>Africa:  Lake Malawi basin (Ref. 2801, 75045).</dc:description>
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	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=2801&amp;speccode=5416'>Lévêque, C. and J. Daget 1984 Cyprinidae. p. 217-342. In J. Daget, J.-P. Gosse and D.F.E. Thys van den Audenaerde (eds.) Check-list of the freshwater fishes of Africa (CLOFFA). ORSTOM, Paris and MRAC, Tervuren. Vol. 1. (Ref. 2801)</reference></dataObject><dataObject>
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	<dc:title>Importance</dc:title>
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	<subject>http://rs.tdwg.org/ontology/voc/SPMInfoItems#DiagnosticDescription</subject>
	<dc:description>fisheries: commercial</dc:description>
	<mediaURL></mediaURL>
	<thumbnailURL></thumbnailURL>
	<location></location>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=4967&amp;speccode=5416'>Eccles, D.H. 1992 FAO species identification sheets for fishery purposes. Field guide to the freshwater fishes of Tanzania. Prepared and published with the support of the United Nations Development Programme (project URT/87/016). FAO, Rome. 145 p. (Ref. 4967)</reference></dataObject><dataObject>
	<dc:identifier>FB-Migration-5416</dc:identifier>
	<dataType>http://purl.org/dc/dcmitype/Text</dataType>
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	<agent homepage='' logoURL='' role='recorder'>Susan M. Luna</agent>
	<dcterms:created>1992-04-20 00:00:00</dcterms:created>
	<dcterms:modified>2024-01-25 00:00:00</dcterms:modified>
	<dc:title></dc:title>
	<dc:language></dc:language>
	<license>http://creativecommons.org/licenses/by-nc/3.0/</license>
	<dc:rights></dc:rights>
	<dcterms:bibliographicCitation></dcterms:bibliographicCitation>
	<dc:source>http://www.fishbase.org/summary/SpeciesSummary.php?id=5416</dc:source>
	<dc:description>Potamodromous. Migrating within streams, migratory in rivers, e.g. Saliminus, Moxostoma, Labeo.  Migrations should be cyclical and predictable and cover more than 100 km.</dc:description>
	<mediaURL></mediaURL>
	<thumbnailURL></thumbnailURL>
	<location></location>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=51243&amp;speccode=5416'>Riede, K. 2004 Global register of migratory species - from global to regional scales. Final Report of the R&amp;D-Projekt 808 05 081. Federal Agency for Nature Conservation, Bonn, Germany. 329 p. (Ref. 51243)</reference></dataObject><dataObject>
	<dc:identifier>FB-TrophicStrategy-5416</dc:identifier>
	<dataType>http://purl.org/dc/dcmitype/Text</dataType>
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	<agent homepage='' logoURL='' role='recorder'>Emmanuel Kaunda</agent>
	<dcterms:created>1992-07-09 00:00:00</dcterms:created>
	<dcterms:modified>2006-01-26 00:00:00</dcterms:modified>
	<dc:title></dc:title>
	<dc:language></dc:language>
	<license>http://creativecommons.org/licenses/by-nc/3.0/</license>
	<dc:rights></dc:rights>
	<dcterms:rightsHolder>WorldFish Center - FishBase</dcterms:rightsHolder>
	<dcterms:bibliographicCitation></dcterms:bibliographicCitation>
	<audience>General public</audience>
	<dc:source>johnstonii</dc:source>
	<subject>http://rs.tdwg.org/ontology/voc/SPMInfoItems#Distribution</subject>
	<dc:description>Inhabits inshore waters and ascends upstream to breed (Ref. 4967).  Found in all types of habitat and occurs in groups of about 25 individuals (Ref. 5595).</dc:description>
	<mediaURL></mediaURL>
	<thumbnailURL></thumbnailURL>
	<location></location>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=4364&amp;speccode=5416'>Jackson, P.B.N. 1961 Check list of the fishes of Nyasaland. The National Museum of Southern Rhodesia. Occasional Papers 3(25B). (Ref. 4364)</reference></dataObject><dataObject>
	<dc:identifier>FB-Size-5416</dc:identifier>
	<dataType>http://purl.org/dc/dcmitype/Text</dataType>
	<mimeType>text/html</mimeType>
	<agent homepage='' logoURL='' role='recorder'>Susan M. Luna</agent>
	<dcterms:created>1992-04-20 00:00:00</dcterms:created>
	<dcterms:modified>2024-01-25 00:00:00</dcterms:modified>
	<dc:title>Max. size</dc:title>
	<dc:language></dc:language>
	<license>http://creativecommons.org/licenses/by-nc/3.0/</license>
	<dc:rights></dc:rights>
	<dcterms:bibliographicCitation></dcterms:bibliographicCitation>
	<dc:source>http://www.fishbase.org/summary/SpeciesSummary.php?id=5416</dc:source>
	<dc:description>50.0 cm SL (male/unsexed; (Ref. 4967))</dc:description>
	<mediaURL></mediaURL>
	<thumbnailURL></thumbnailURL>
	<location></location>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=4967&amp;speccode=5416'>Eccles, D.H. 1992 FAO species identification sheets for fishery purposes. Field guide to the freshwater fishes of Tanzania. Prepared and published with the support of the United Nations Development Programme (project URT/87/016). FAO, Rome. 145 p. (Ref. 4967)</reference></dataObject><dataObject>
	<dc:identifier>FB-Threats-2</dc:identifier>
	<dataType>http://purl.org/dc/dcmitype/Text</dataType>
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	<agent homepage='' logoURL='' role='recorder'>Belen Acosta</agent>
	<dcterms:created>1990-10-18 00:00:00</dcterms:created>
	<dcterms:modified>2024-03-07 00:00:00</dcterms:modified>
	<dc:title></dc:title>
	<dc:language></dc:language>
	<license>http://creativecommons.org/licenses/by-nc/3.0/</license>
	<dc:rights></dc:rights>
	<dcterms:rightsHolder>WorldFish Center - FishBase</dcterms:rightsHolder>
	<dcterms:bibliographicCitation></dcterms:bibliographicCitation>
	<audience>General public</audience>
	<dc:source>http://www.fishbase.org/summary/SpeciesSummary.php?id=2</dc:source>
	<subject>http://rs.tdwg.org/ontology/voc/SPMInfoItems#Size</subject>
	<dc:description>Least Concern (LC) , IUCN Grouper and Wrasse Specialist Group</dc:description>
	<mediaURL></mediaURL>
	<thumbnailURL></thumbnailURL>
	<location></location>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=57073'>IUCN 2006 2006 IUCN red list of threatened species. www.iucnredlist.org. Downloaded July 2006.</reference></dataObject><dataObject>
	<dc:identifier>FB-Habitat-5416</dc:identifier>
	<dataType>http://purl.org/dc/dcmitype/Text</dataType>
	<mimeType>text/html</mimeType>
	<agent homepage='' logoURL='' role='recorder'>Susan M. Luna</agent>
	<dcterms:created>1992-04-20 00:00:00</dcterms:created>
	<dcterms:modified>2024-01-25 00:00:00</dcterms:modified>
	<dc:title>Environment</dc:title>
	<dc:language></dc:language>
	<license>http://creativecommons.org/licenses/by-nc/3.0/</license>
	<dc:rights></dc:rights>
	<dcterms:rightsHolder>WorldFish Center - FishBase</dcterms:rightsHolder>
	<dcterms:bibliographicCitation></dcterms:bibliographicCitation>
	<audience>General public</audience>
	<dc:source>http://www.fishbase.org/summary/SpeciesSummary.php?id=5416</dc:source>
	<subject>http://rs.tdwg.org/ontology/voc/SPMInfoItems#Threats</subject>
	<dc:description>benthopelagic; potamodromous (Ref. 51243); freshwater</dc:description>
	<mediaURL></mediaURL>
	<thumbnailURL></thumbnailURL>
	<location></location>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=51243&amp;speccode=5416'>Riede, K. 2004 Global register of migratory species - from global to regional scales. Final Report of the R&amp;D-Projekt 808 05 081. Federal Agency for Nature Conservation, Bonn, Germany. 329 p. (Ref. 51243)</reference></dataObject>
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