FB-5015 http://www.fishbase.org/summary/SpeciesSummary.php?id=5015 Animalia Chordata Elasmobranchii Rajiformes Rajidae Raja Raja straeleni Poll, 1951 Peixe-areia Raia-pontuada-africana Rog Vals doringrug-rog Biscuit skate False thornback skate Skate Spotted skate Präänikrai Asifome Raie tachetée Tantra sese Raya manchada Biscuitrog Tantre 斯氏鰩 斯氏鳐 Raja maderensis (non Lowe, 1838) 1992-02-11 2024-01-23 Breder, C.M. and D.E. Rosen0 Modes of reproduction in fishes. T.F.H. Publications, Neptune City, New Jersey. 941 p. (Ref. 205) Poll, M.0 Expédition océanographique belge dans les eaux côtières africaines de l'Atlantique Sud (1948-1949). Résultats scientifiques. Volume IV, fascicule 1. Poissons. I. Généralités. II. Sélaciens et chimères. Institut Royal des Sciences Naturelles de Belgique. 154 pp. (Ref. 377) Schneider, W.0 FAO species identification sheets for fishery purposes. Field guide to the commercial marine resources of the Gulf of Guinea. Prepared and published with the support of the FAO Regional Office for Africa. Rome: FAO. 268 p. (Ref. 2683) Hulley, P.A.0 Rajidae. p. 115-127. In M.M. Smith and P.C. Heemstra (eds.) Smiths' sea fishes. Springer-Verlag, Berlin. (Ref. 2708) Stehmann, M.0 Rajidae. p. 29-50. In J.C. Quero, J.C. Hureau, C. Karrer, A. Post and L. Saldanha (eds.) Check-list of the fishes of the eastern tropical Atlantic. Junta Nacional de Investigaçao Cientifica e Tecnológica, Lisbon, Portugal. Vol. 1. (Ref. 4426) Hureau, J.-C.0 La base de données GICIM: Gestion informatisée des collections ichthyologiques du Muséum. p. 225-227. In Atlas Préliminaire des Poissons d'Eau Douce de France. Conseil Supérieur de la Pêche, Ministère de l'Environnement, CEMAGREF et Muséum national d'Histoire naturelle, Paris. (Ref. 4517) Maigret, J. and B. Ly0 Les poissons de mer de Mauritanie. Science Nat., Compiègne. 213 p. (Ref. 5377) Compagno, L.J.V., D.A. Ebert and M.J. Smale0 Guide to the sharks and rays of southern Africa. New Holland (Publ.) Ltd., London. 158 p. (Ref. 5578) Fritzsch, B. and P. Moller0 A history of electroreception. p. 39-55. In P. Moller (ed.) Electric fishes:history and behavior. Fish and Fisheries Series 17. Chapman & Hall, London. (Ref. 10311) Stehmann, M.0 First and new records of skates (Chondrichthyes, Rajiformes, Rajidae) from the West African continental slope (Morocco to South Africa), with descriptions of two new species. Arch. Fish. Mar. Res. 43(1):1-119. (Ref. 11284) Sanches, J.G.0 Catálogo dos principais peixes marinhos da República de Guiné-Bissau. Publ. Avuls. Inst. Nac. Invest. Pescas 16:429 p. (Ref. 26999) Bianchi, G., K.E. Carpenter, J.-P. Roux, F.J. Molloy, D. Boyer and H.J. Boyer0 FAO species identification guide for fishery purposes. Field guide to the living marine resources of Namibia. FAO, Rome. 265p. (Ref. 27121) McEachran, J.D. and K.A. Dunn0 Phylogenetic analysis of skates, a morphologically conservative clade of elasmobranchs (Chondrichthyes: Rajidae). Copeia 1998(2):271-290. (Ref. 27314) Macpherson, E. and B.A. Roel0 Trophic relationships in the demersal fish community off Namibia. S. Afr. J. Mar. Sci. 5:585-596. (Ref. 27637) Wu, H.L., K.-T. Shao and C.F. Lai (eds.)0 Latin-Chinese dictionary of fishes names. The Sueichan Press, Taiwan. 1028 p. (Ref. 31517) Anonymous0 Fish collection database of the Natural History Museum, London (formerly British Museum of Natural History (BMNH)). Natural History Museum, London (formerly British Museum of Natural History (BMNH)). (Ref. 31982) Compagno, L.J.V.0 Checklist of living elasmobranchs. p. 471-498. In W.C. Hamlett (ed.) Sharks, skates, and rays: the biology of elasmobranch fishes. Johns Hopkins University Press, Maryland. (Ref. 35766) Anonymous0 Fish collection database of the J.L.B. Smith Institute of Ichthyology, Grahamstown, South Africa. J.L.B. Smith Institute of Ichthyology, Grahamstown, South Africa. (Ref. 36670) Bianchi, G., K.E. Carpenter, J.-P. Roux, F.J. Molloy, D. Boyer and H.J. Boyer0 FAO species identification guide for fishery purposes. Field guide to the living marine resources of Namibia. FAO, Rome. 265 p. (Ref. 36731) Anonymous0 Fish collection database of the National Museum of Natural History (Smithsonian Institution). Smithsonian Institution-Division of Fishes. (Ref. 38732) Haedrich, R.L. and N.R. Merrett0 Summary atlas of deep-living demersal fishes in the North Atlantic Basin. J. Nat. Hist. 22:1325-1362. (Ref. 40643) Anonymous0 Fish collection database of the Zoological Museum, University of Copenhagen. Zoological Museum, University of Copenhagen. (Ref. 40919) Bor, P.H.F.0 Nederlandse naamlijst van de recente haaien en roggen (Chondrichthyes: Elasmobranchii) van de wereld. World Wide Web electronic publication www.rajidae.tmfweb.nl, version (05/2002). (Ref. 43454) Dulvy, N.K. and J.D. Reynolds0 Evolutionary transitions among egg-laying, live-bearing and maternal inputs in sharks and rays. Proc. R. Soc. Lond., Ser. B: Biol. Sci. 264:1309-1315. (Ref. 50449) Okeyo, D.O.0 Fishes of Namibia. Unpublished compilation of D.O. Okeyo. (Ref. 54854) Mohamed Fall, K.O.0 Fishes of Mauritania. Unpublished compilation of K.O. Mohamed Fall. (Ref. 55783) Chinese Academy of Fishery Sciences0 Chinese aquatic germplasm resources database. http://zzzy.fishinfo.cn (Ref. 58108) Macpherson, E.0 Influence of geographical distribution, body size and diet on population density of benthic fishes off Namibia (South West Africa). Mar. Ecol. Prog. Ser. 50:295-299. (Ref. 58748) FAO-FIES0 Aquatic Sciences and Fisheries Information System (ASFIS) species list. Retrieved from http://www.fao.org/fishery/collection/asfis, 29 April 2008. (Ref. 75549) Ebert, D.A., L.J.V. Compagno and P.D. Cowley0 Aspects of the reproductive biology of skates (Chondrichthyes: Rajiformes: Rajoidei) from southern Africa. ICES Journal of Marine Science 65(1):81-102. (Ref. 80491) FAO-FIES0 Aquatic Sciences and Fisheries Information System (ASFIS) species list. Retrieved from http://www.fao.org/fishery/collection/asfis/en, March 2010. (Ref. 83882) FAO-FIES0 Aquatic Sciences and Fisheries Information System (ASFIS) species list. Retrieved from http://www.fao.org/fishery/collection/asfis/en, March 2012. (Ref. 90062) FAO-FIES0 Aquatic Sciences and Fisheries Information System (ASFIS) species list. Retrieved from http://www.fao.org/fishery/collection/asfis/en, April 2014. (Ref. 95632) FAO-FIES0 Aquatic Sciences and Fisheries Information System (ASFIS) species list. Retrieved from http://www.fao.org/fishery/collection/asfis/en, [accessed 13/04/2015]. (Ref. 101110) eElurikkus0 eElurikkus koondab ühte portaali Eesti eluslooduse andmed. http://elurikkus.ut.ee/ (Ref. 105471) FAO-FIES0 Aquatic Sciences and Fisheries Information System (ASFIS) species list. Retrieved from http://www.fao.org/fishery/collection/asfis/en (accessed 08/06/2017). (Ref. 115257) FAO-FIES0 Aquatic Sciences and Fisheries Information System (ASFIS) species list. Retrieved from htttps://www.fao.org/fishery/collection/asfis/en [accessed May, 2018] (Ref. 118272) Beolens, B., M. Grayson and M. Watkins0 Eponym dictionary of Fishes. Dunbeath, Caithness (Scotland, UK): Whittles Publishing Limited, 1300 p. (Ref. 128868) Kwei, E.A. and D.W. Ofori-Adu0 Fishes in the coastal waters of Ghana. Comm. 2. Tema-Ghana. 124p. (Ref. 130891) Ferragut-Perello, F., S. Ramirez-Amaro, A.C. Tsikliras, N. Petit-Marty, D. Dimarchopoulou0 Exploitation and conservation status of the thornback ray (<i>Raja clavata</i>) in the Balearic Islands (Western Mediterranean). Fishes 8(2):117. (Ref. 132125) FB-GeneralDescr-5015 http://purl.org/dc/dcmitype/Text text/html Susan M. Luna 1992-02-11 00:00:00 2024-01-23 00:00:00 Biology http://creativecommons.org/licenses/by-nc/3.0/ WorldFish Center - FishBase General public http://www.fishbase.org/summary/SpeciesSummary.php?id=5015 http://rs.tdwg.org/ontology/voc/SPMInfoItems#Habitat Found in shallow enclosed bays and on the slope (Ref. 5578). Feeds on invertebrates, bony fishes and fish offal. Oviparous (Ref. 50449). Eggs have horn-like projections on the shell (Ref. 205). Caught by shore and ski-boat anglers and by hake trawlers. Pectoral fins sold as 'skate wings' (Ref. 5578). Undergoes regular migration, towards the coast during the warm season and towards deep waters in winter (Ref. 5377). McEachran, J.D. and K.A. Dunn 1998 Phylogenetic analysis of skates, a morphologically conservative clade of elasmobranchs (Chondrichthyes: Rajidae). Copeia 1998(2):271-290. (Ref. 27314) FB-DiagnosticDescr-5015 http://purl.org/dc/dcmitype/Text text/html Cristina V. Garilao 1996-11-25 00:00:00 2017-11-15 00:00:00 http://creativecommons.org/licenses/by-nc/3.0/ WorldFish Center - FishBase General public http://www.fishbase.org/summary/SpeciesSummary.php?id=5015 http://rs.tdwg.org/ontology/voc/SPMInfoItems#Distribution A black-spotted skate with a blunt, angular snout, a long, stout tail, and a broad angular disc; strong thorns on upper surface of disc and tail at all stages, sometimes on underside in large individuals (Ref. 5578). Brown to grey dorsally, with small to large black spots, whorls and blotches; many with gold and black, traversely oblong eyespot on each pectoral base; white ventrally with no dark pores (Ref. 5578). Juveniles have no dark spots above (Ref. 2708). Compagno, L.J.V., D.A. Ebert and M.J. Smale 1989 Guide to the sharks and rays of southern Africa. New Holland (Publ.) Ltd., London. 158 p. (Ref. 5578) FB-Distribution-5015 http://purl.org/dc/dcmitype/Text text/html Susan M. Luna 1992-02-11 00:00:00 2022-06-22 00:00:00 http://creativecommons.org/licenses/by-nc/3.0/ http://www.fishbase.org/summary/SpeciesSummary.php?id=5015 Eastern Atlantic: Rio de Oro, Western Sahara to South Africa. Stehmann, M. 1995 First and new records of skates (Chondrichthyes, Rajiformes, Rajidae) from the West African continental slope (Morocco to South Africa), with descriptions of two new species. Arch. Fish. Mar. Res. 43(1):1-119. (Ref. 11284) FB-Uses-5015 http://purl.org/dc/dcmitype/Text text/html Susan M. Luna 1992-02-11 00:00:00 2024-01-23 00:00:00 Importance http://creativecommons.org/licenses/by-nc/3.0/ WorldFish Center - FishBase General public http://www.fishbase.org/summary/SpeciesSummary.php?id=5015 http://rs.tdwg.org/ontology/voc/SPMInfoItems#Size fisheries: commercial; gamefish: yes Schneider, W. 1990 FAO species identification sheets for fishery purposes. Field guide to the commercial marine resources of the Gulf of Guinea. Prepared and published with the support of the FAO Regional Office for Africa. Rome: FAO. 268 p. (Ref. 2683) Compagno, L.J.V., D.A. Ebert and M.J. Smale 1989 Guide to the sharks and rays of southern Africa. New Holland (Publ.) Ltd., London. 158 p. (Ref. 5578) FB-Migration-5015 http://purl.org/dc/dcmitype/Text text/html Susan M. Luna 1992-02-11 00:00:00 2024-01-23 00:00:00 http://creativecommons.org/licenses/by-nc/3.0/ WorldFish Center - FishBase General public http://www.fishbase.org/summary/SpeciesSummary.php?id=5015 http://rs.tdwg.org/ontology/voc/SPMInfoItems#Threats Oceanodromous. Migrating within oceans typically between spawning and different feeding areas, as tunas do. Migrations should be cyclical and predictable and cover more than 100 km. FB-LifeCycle-5015 http://purl.org/dc/dcmitype/Text text/html Susan M. Luna 1992-02-11 00:00:00 2024-01-23 00:00:00 http://creativecommons.org/licenses/by-nc/3.0/ WorldFish Center - FishBase General public http://www.fishbase.org//Reproduction/FishReproSummary.php?ID=5015&StockCode=5249 http://rs.tdwg.org/ontology/voc/SPMInfoItems#Habitat Oviparous, paired eggs are laid. Embryos feed solely on yolk (Ref. 50449). Breder, C.M. and D.E. Rosen 1966 Modes of reproduction in fishes. T.F.H. Publications, Neptune City, New Jersey. 941 p. (Ref. 205) FB-TrophicStrategy-5015 http://purl.org/dc/dcmitype/Text text/html Pascualita Sa-a 1999-12-07 00:00:00 2007-08-28 00:00:00 http://creativecommons.org/licenses/by-nc/3.0/ WorldFish Center - FishBase General public straeleni http://rs.tdwg.org/ontology/voc/SPMInfoItems#Distribution Found in shallow enclosed bays and on the slope (Ref. 5578). Feeds on invertebrates, bony fishes and fish offal. Stehmann, M. 1995 First and new records of skates (Chondrichthyes, Rajiformes, Rajidae) from the West African continental slope (Morocco to South Africa), with descriptions of two new species. Arch. Fish. Mar. Res. 43(1):1-119. (Ref. 11284) FB-Size-5015 http://purl.org/dc/dcmitype/Text text/html Susan M. Luna 1992-02-11 00:00:00 2024-01-23 00:00:00 Max. size http://creativecommons.org/licenses/by-nc/3.0/ http://www.fishbase.org/summary/SpeciesSummary.php?id=5015 70.0 cm TL (male/unsexed; (Ref. 26999)); 67 cm TL (female); max. published weight: 1,385 g (Ref. 377); max. published weight: 1,685.0 g Sanches, J.G. 1991 Catálogo dos principais peixes marinhos da República de Guiné-Bissau. Publ. Avuls. Inst. Nac. Invest. Pescas 16:429 p. (Ref. 26999) Poll, M. 1951 Expédition océanographique belge dans les eaux côtières africaines de l'Atlantique Sud (1948-1949). Résultats scientifiques. Volume IV, fascicule 1. Poissons. I. Généralités. II. Sélaciens et chimères. Institut Royal des Sciences Naturelles de Belgique. 154 pp. (Ref. 377) FB-Threats-2 http://purl.org/dc/dcmitype/Text text/html Belen Acosta 1990-10-18 00:00:00 2024-03-07 00:00:00 http://creativecommons.org/licenses/by-nc/3.0/ http://www.fishbase.org/summary/SpeciesSummary.php?id=2 Least Concern (LC) , IUCN Grouper and Wrasse Specialist Group IUCN 2006 2006 IUCN red list of threatened species. www.iucnredlist.org. Downloaded July 2006. FB-Habitat-5015 http://purl.org/dc/dcmitype/Text text/html Susan M. Luna 1992-02-11 00:00:00 2024-01-23 00:00:00 Environment http://creativecommons.org/licenses/by-nc/3.0/ WorldFish Center - FishBase General public http://www.fishbase.org/summary/SpeciesSummary.php?id=5015 http://rs.tdwg.org/ontology/voc/SPMInfoItems#Threats demersal; oceanodromous; marine; depth range 80 - 800 m (Ref. 11284), usually 100 - 300 m (Ref. 26999) Stehmann, M. 1995 First and new records of skates (Chondrichthyes, Rajiformes, Rajidae) from the West African continental slope (Morocco to South Africa), with descriptions of two new species. Arch. Fish. Mar. Res. 43(1):1-119. (Ref. 11284) Sanches, J.G. 1991 Catálogo dos principais peixes marinhos da República de Guiné-Bissau. Publ. Avuls. Inst. Nac. Invest. Pescas 16:429 p. (Ref. 26999)