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	<dc:identifier>FB-48130</dc:identifier>
	<dc:source>http://www.fishbase.org/summary/SpeciesSummary.php?id=48130</dc:source>
	<dwc:Kingdom>Animalia</dwc:Kingdom>
	<dwc:Phylum>Chordata</dwc:Phylum>
	<dwc:Class>Teleostei</dwc:Class>
	<dwc:Order>Gymnotiformes</dwc:Order>
	<dwc:Family>Gymnotidae</dwc:Family>
	<dwc:Genus>Gymnotus</dwc:Genus>
	<dwc:ScientificName>Gymnotus pantherinus (Steindachner, 1908)</dwc:ScientificName>
	<commonName xml:lang='da'>Valaam-helt</commonName>
	<commonName xml:lang='de'>Schalsee Maräne</commonName>
	<commonName xml:lang='de'>Selenter Maräne</commonName>
	<commonName xml:lang='en'>Pool whitefish</commonName>
	<commonName xml:lang='en'>Valaam whitefish</commonName>
	<commonName xml:lang='en'>Valaamka</commonName>
	<commonName xml:lang='ru'>Kryazhevoi sig</commonName>
	<commonName xml:lang='ru'>Valaamka</commonName>
	<commonName xml:lang='ru'>Yamnyi sig</commonName>
	<commonName xml:lang='ru'>Zobatyi sig</commonName>
	<commonName xml:lang='ru'>валаамка</commonName>
	<commonName xml:lang='sv'>Sandsik</commonName>
	<synonym relationship='misapplied name'>Coregonus baerii (non Kessler, 1864)</synonym>
	<synonym relationship='synonym'>Coregonus gracilis Günther, 1866</synonym>
	<synonym relationship='synonym'>Coregonus holsatus anarensis Järvi, 1928</synonym>
	<synonym relationship='synonym'>Coregonus holsatus scallensis Thienemann, 1921</synonym>
	<synonym relationship='synonym'>Coregonus holsatus suecica Thienemann, 1921</synonym>
	<synonym relationship='synonym'>Coregonus lavaretus holsatus f. okoniensis Kulmatycki, 1926</synonym>
	<synonym relationship='ambiguous synonym'>Coregonus lavaretus ludoga Polyakov, 1874</synonym>
	<synonym relationship='synonym'>Coregonus lavaretus okoniensis Kulmatycki, 1926</synonym>
	<synonym relationship='ambiguous synonym'>Coregonus lavaretus poljakovi Pravdin, 1931</synonym>
	<synonym relationship='ambiguous synonym'>Coregonus lavaretus poljakowi Pravdin, 1931</synonym>
	<synonym relationship='synonym'>Coregonus lavaretus polonica Kulmatycki, 1926</synonym>
	<synonym relationship='synonym'>Coregonus lavaretus widegreni Malmgren, 1863</synonym>
	<synonym relationship='ambiguous synonym'>Coregonus ludoga Polyakov, 1874</synonym>
	<synonym relationship='ambiguous synonym'>Coregonus ludoga swirensis Polyakov, 1874</synonym>
	<synonym relationship='ambiguous synonym'>Coregonus ludoga verchoswirka Polyakov, 1874</synonym>
	<synonym relationship='misapplied name'>Coregonus maraena holsata (non Thienemann, 1916)</synonym>
	<synonym relationship='ambiguous synonym'>Coregonus tscholmugensis Danilewsky, 1873</synonym>
	<synonym relationship='misapplied name'>Coregonus widegreni ludoga (non Berg, 1916)</synonym>
	<dcterms:created>1998-04-30</dcterms:created>
	<dcterms:modified>2015-02-10</dcterms:modified>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=205&amp;speccode=48137'>Breder, C.M. and D.E. Rosen0 Modes of reproduction in fishes. T.F.H. Publications, Neptune City, New Jersey. 941 p. (Ref. 205)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=593&amp;speccode=48137'>Berg, L.S.0 Freshwater fishes of the U.S.S.R. and adjacent countries. volume 1, 4th edition. Israel Program for Scientific Translations Ltd, Jerusalem. (Russian version published 1948). (Ref. 593)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=6094&amp;speccode=48137'>Chiba, K., Y. Taki, K. Sakai and Y. Oozeki0 Present status of aquatic organisms introduced into Japan. p. 63-70. In S.S. De Silva (ed.) Exotic aquatic organisms in Asia. Proceedings of the Workshop on Introduction of Exotic Aquatic Organisms in Asia. Spec. Publ. Asian Fish. Soc. 3, 154 p. (Ref. 6094)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=6275&amp;speccode=48137'>Huusko, A.0 Kuusinkijoen kalatalousselvitys. (Study on the fisheries of the River Kuusinkijoki). Riista-ja kalatalouden tutkimuslaitos. Kalatutkimuksia 14. Helsinki, Finland. (Publication of the Finnish Game and Fisheries Research Institute). (in Finnish). (Ref. 6275)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=13696&amp;speccode=48137'>Kottelat, M.0 European freshwater fishes. An heuristic checklist of the freshwater fishes of Europe (exclusive of former USSR), with an introduction for non-systematists and comments on nomenclature and conservation. Biologia, Bratislava, 52/Suppl. 5:1-271. (Ref. 13696)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=30184&amp;speccode=48137'>Arkhipchuk, V.V.0 Chromosome database. Database of Dr. Victor Arkhipchuk. (Ref. 30184)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=30402&amp;speccode=48137'>Kullander, S.O.0 Swedish fishes: checklist of Swedish fishes. Swedish Museum of Natural History. http://www.nrm.se/ve/pisces/allfish.shtml.en, 25 May 1999. (Ref. 30402)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=31670&amp;speccode=48137'>Lehtonen, H.0 Biology and stock assessments of coregonids by the Baltic coast of Finland. Finnish Fish. Res. 3:31-83. (Ref. 31670)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=39901&amp;speccode=48137'>Wikgren, B.-J.0 Resultaten av sikmärkningar inom Aland och vid Luvia. Husš Biol. Stat. Medd. 3:1-26. (Ref. 39901)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=45022&amp;speccode=48137'>Bogutskaya, N.G. and A.M. Naseka0 An overview of nonindigenous fishes in inland waters of Russia. Proc. Zool. Inst. Russ. Acad. Sci. 296:21-30. (Ref. 45022)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=45043&amp;speccode=48137'>Bogutskaya, N.G.0 Common names of Russian freshwater fishes. Spreadsheet with cyrillic names made available for inclusion in FishBase. Unpublished. (Ref. 45043)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=51471&amp;speccode=48137'>Carl, H.0 Danish fish names. Zoological Museum of Copenhagen. Unpublished. (Ref. 51471)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=56506&amp;speccode=48137'>Morozova, P.N.0 Fisheries and biological characteristics of whitefishes of Ladoga Lake. Izv. VNIORH 38:95-109. (Ref. 56506)</reference>
	<reference isbn='978-2-8399-0298-4' url='http://www.fishbase.org/references/FBRefSummary.php?id=59043&amp;speccode=48137'>Kottelat, M. and J. Freyhof0 Handbook of European freshwater fishes. Publications Kottelat, Cornol and Freyhof, Berlin. 646 pp. (Ref. 59043)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=74657&amp;speccode=48137'>Bartley, D.M. (comp./ed.)0 DIAS. Introduced species in fisheries and aquaculture: information for responsible use and control. CD-ROM. Rome (Italy): FAO. (Ref. 74657)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=75549&amp;speccode=48137'>FAO-FIES0 Aquatic Sciences and Fisheries Information System (ASFIS) species list. Retrieved from http://www.fao.org/fishery/collection/asfis, 29 April 2008. (Ref. 75549)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=78639&amp;speccode=48137'>Kottelat, M., N.G. Bogutskaya and J. Freyhof0 On the migratory Black Sea lamprey and the nomenclature of the ludoga, Peipsi and ripus whitefishes (Agnatha: Petromyzontidae; Teleostei: Coregonidae). Zoosyst. Rossica 14(1):181-186. (Ref. 78639)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=83882&amp;speccode=48137'>FAO-FIES0 Aquatic Sciences and Fisheries Information System (ASFIS) species list. Retrieved from http://www.fao.org/fishery/collection/asfis/en, March 2010. (Ref. 83882)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=89336&amp;speccode=48137'>Eschmeyer, W.N. and Fricke R. (eds.)0 Catalog of fishes. Updated internet version of 12 January 2012. Catalog databases of CAS cited in FishBase (website). (Ref. 89336)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=90062&amp;speccode=48137'>FAO-FIES0 Aquatic Sciences and Fisheries Information System (ASFIS) species list. Retrieved from http://www.fao.org/fishery/collection/asfis/en, March 2012. (Ref. 90062)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=95632&amp;speccode=48137'>FAO-FIES0 Aquatic Sciences and Fisheries Information System (ASFIS) species list. Retrieved from http://www.fao.org/fishery/collection/asfis/en, April 2014. (Ref. 95632)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=101110&amp;speccode=48137'>FAO-FIES0 Aquatic Sciences and Fisheries Information System (ASFIS) species list. Retrieved from http://www.fao.org/fishery/collection/asfis/en, [accessed 13/04/2015]. (Ref. 101110)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=115257&amp;speccode=48137'>FAO-FIES0 Aquatic Sciences and Fisheries Information System (ASFIS) species list. Retrieved from http://www.fao.org/fishery/collection/asfis/en (accessed 08/06/2017). (Ref. 115257)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=116654&amp;speccode=48137'>Saks, L.0 Estonian Common Names, Rank 2. Personal communication (Ref. 116654)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=118272&amp;speccode=48137'>FAO-FIES0 Aquatic Sciences and Fisheries Information System (ASFIS) species list. Retrieved from htttps://www.fao.org/fishery/collection/asfis/en [accessed May, 2018] (Ref. 118272)</reference>
	<reference isbn='1849954984' url='http://www.fishbase.org/references/FBRefSummary.php?id=128868&amp;speccode=48137'>Beolens, B., M. Grayson and M. Watkins0 Eponym dictionary of Fishes. 	Dunbeath, Caithness (Scotland, UK): Whittles Publishing Limited, 1300 p. (Ref. 128868)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=130435&amp;speccode=48137'>IUCN0 The IUCN Red List of Threatened Species. 1964 - onwards. https://www.iucnredlist.org/ (Ref. 130435)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=138280&amp;speccode=48137'>Corush, J.B.0 Evolutionary patterns of diadromy in fishes: more than a transitional state between marine and freshwater. BMC Evol. Biol. 19(168):13 p. (Ref. 138280)</reference><dataObject>
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	<agent homepage='' logoURL='' role='recorder'>Pascualita Sa-a</agent>
	<dcterms:created>1998-05-01 00:00:00</dcterms:created>
	<dcterms:modified>2025-11-18 00:00:00</dcterms:modified>
	<dc:title>Biology</dc:title>
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	<dc:description>Recorded from coastal waters at sea. Lacustrine, at depths greater than 30 m, in Lake Ladoga commonly at 50-150 m (Ref. 59043). Specimens measuring 36-40 cm are more than 5 years old.  Mature individuals spawn for the first time at 4-6 years (Ref. 59043) from the end of November to December (Ref. 593).  Oviparous (Ref. 205).  Eggs are buried in unguarded nests (Ref. 205). Adults feed on benthic invertebrates, especially gammarids, molluscs and insect larvae (Ref. 59043).</dc:description>
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	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=13696&amp;speccode=48137'>Kottelat, M. 1997 European freshwater fishes. An heuristic checklist of the freshwater fishes of Europe (exclusive of former USSR), with an introduction for non-systematists and comments on nomenclature and conservation. Biologia, Bratislava, 52/Suppl. 5:1-271. (Ref. 13696)</reference></dataObject><dataObject>
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	<dcterms:created>1998-05-01 00:00:00</dcterms:created>
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	<dc:description>Europe:  Baltic: Bothnian Gulf, Gotland (missing along Swedish coast southward of Stockholm); southern Baltic: from Germany to Gulf of Finland; several landlocked populations in Sweden (Lakes Vänern, Vättern, Locknesjön, Storvindeln, Storlaisan, Skalka-Parkijaur, Arjeplog) and Russia (Neva drainage, Lakes Ladoga and Onega).</dc:description>
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	<reference isbn='978-2-8399-0298-4' url='http://www.fishbase.org/references/FBRefSummary.php?id=59043&amp;speccode=48137'>Kottelat, M. and J. Freyhof 2007 Handbook of European freshwater fishes. Publications Kottelat, Cornol and Freyhof, Berlin. 646 pp. (Ref. 59043)</reference></dataObject><dataObject>
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	<dcterms:created>1998-05-01 00:00:00</dcterms:created>
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	<dc:description>Anadromous. Fish that ascend rivers to spawn, as salmon and hilsa do. Sub-division of diadromous.    Migrations should be cyclical and predictable and cover more than 100 km.</dc:description>
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	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=138280&amp;speccode=48137'>Corush, J.B. 2019 Evolutionary patterns of diadromy in fishes: more than a transitional state between marine and freshwater. BMC Evol. Biol. 19(168):13 p. (Ref. 138280)</reference></dataObject><dataObject>
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	<agent homepage='' logoURL='' role='recorder'>Susan M. Luna</agent>
	<dcterms:created>1998-05-01 00:00:00</dcterms:created>
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	<dc:source>http://www.fishbase.org//Reproduction/FishReproSummary.php?ID=48137&amp;StockCode=40417</dc:source>
	<dc:description>Distinct pairing (Ref. 205).</dc:description>
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	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=205&amp;speccode=48137'>Breder, C.M. and D.E. Rosen 1966 Modes of reproduction in fishes. T.F.H. Publications, Neptune City, New Jersey. 941 p. (Ref. 205)</reference></dataObject><dataObject>
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	<dc:title>Max. size</dc:title>
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	<dc:description>55.0 cm SL (male/unsexed; (Ref. 59043)); max. published weight: 2,000 g (Ref. 593); max. reported age: 20 years (Ref. 59043)</dc:description>
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	<reference isbn='978-2-8399-0298-4' url='http://www.fishbase.org/references/FBRefSummary.php?id=59043&amp;speccode=48137'>Kottelat, M. and J. Freyhof 2007 Handbook of European freshwater fishes. Publications Kottelat, Cornol and Freyhof, Berlin. 646 pp. (Ref. 59043)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=593&amp;speccode=48137'>Berg, L.S. 1962 Freshwater fishes of the U.S.S.R. and adjacent countries. volume 1, 4th edition. Israel Program for Scientific Translations Ltd, Jerusalem. (Russian version published 1948). (Ref. 593)</reference>
	<reference isbn='978-2-8399-0298-4' url='http://www.fishbase.org/references/FBRefSummary.php?id=59043&amp;speccode=48137'>Kottelat, M. and J. Freyhof 2007 Handbook of European freshwater fishes. Publications Kottelat, Cornol and Freyhof, Berlin. 646 pp. (Ref. 59043)</reference></dataObject><dataObject>
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	<dc:description>Least Concern (LC) , IUCN Grouper and Wrasse Specialist Group</dc:description>
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	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=57073'>IUCN 2006 2006 IUCN red list of threatened species. www.iucnredlist.org. Downloaded July 2006.</reference></dataObject><dataObject>
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	<subject>http://rs.tdwg.org/ontology/voc/SPMInfoItems#Habitat</subject>
	<dc:description>demersal; anadromous (Ref. 138280); freshwater; depth range 30 - 150 m (Ref. 59043)</dc:description>
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	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=138280&amp;speccode=48137'>Corush, J.B. 2019 Evolutionary patterns of diadromy in fishes: more than a transitional state between marine and freshwater. BMC Evol. Biol. 19(168):13 p. (Ref. 138280)</reference>
	<reference isbn='978-2-8399-0298-4' url='http://www.fishbase.org/references/FBRefSummary.php?id=59043&amp;speccode=48137'>Kottelat, M. and J. Freyhof 2007 Handbook of European freshwater fishes. Publications Kottelat, Cornol and Freyhof, Berlin. 646 pp. (Ref. 59043)</reference></dataObject>
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