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	<dc:identifier>FB-48083</dc:identifier>
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	<dwc:Kingdom>Animalia</dwc:Kingdom>
	<dwc:Phylum>Chordata</dwc:Phylum>
	<dwc:Class>Teleostei</dwc:Class>
	<dwc:Order>Perciformes/Cottoidei</dwc:Order>
	<dwc:Family>Cottidae</dwc:Family>
	<dwc:Genus>Limnocottus</dwc:Genus>
	<dwc:ScientificName>Limnocottus pallidus Taliev, 1948</dwc:ScientificName>
	<commonName xml:lang='cs'>Sleď tanajský</commonName>
	<commonName xml:lang='en'>Azov shad</commonName>
	<commonName xml:lang='en'>Black Sea shad</commonName>
	<commonName xml:lang='en'>Paeostom shad</commonName>
	<commonName xml:lang='et'>Tanaisi aloosa</commonName>
	<commonName xml:lang='ru'>Azovskii puzanok</commonName>
	<commonName xml:lang='ru'>Azowskyi puzanok</commonName>
	<commonName xml:lang='ru'>Dunayskyi puzanok</commonName>
	<commonName xml:lang='ru'>Palaeostomskii puzanok</commonName>
	<commonName xml:lang='ru'>Paliastomskyi puzanok</commonName>
	<commonName xml:lang='ru'>азовская сельдь</commonName>
	<commonName xml:lang='tr'>Ringa balığı</commonName>
	<commonName xml:lang='tr'>Tirsi</commonName>
	<commonName xml:lang='tr'>Tirsi balığı</commonName>
	<commonName xml:lang=''>Azovska čepa</commonName>
	<commonName xml:lang=''>Rizeafca</commonName>
	<commonName xml:lang=''>Rizeafcă</commonName>
	<synonym relationship='synonym'>Alosa bulgarica Drensky, 1934</synonym>
	<synonym relationship='synonym'>Alosa caspia nordmanni Antipa, 1904</synonym>
	<synonym relationship='synonym'>Alosa caspia tanaica (Grimm, 1901)</synonym>
	<synonym relationship='ambiguous synonym'>Alosa caspia tanaica asovi Pavlov, 1959</synonym>
	<synonym relationship='synonym'>Alosa nordmanni Antipa, 1904</synonym>
	<synonym relationship='synonym'>Caspialosa caspia palaeostomi Sadowsky, 1934</synonym>
	<synonym relationship='synonym'>Caspialosa caspia tanaica (Grimm, 1901)</synonym>
	<synonym relationship='synonym'>Caspialosa palaeostomica Sadowsky, 1934</synonym>
	<synonym relationship='ambiguous synonym'>Caspialosa tanaica etemi Battalgil, 1941</synonym>
	<synonym relationship='synonym'>Caspialosa tanaica palaeostomi Sadowsky, 1934</synonym>
	<synonym relationship='ambiguous synonym'>Caspiolosa tanaica etemi Battalgil, 1941</synonym>
	<synonym relationship='synonym'>Clupea tanaica Grimm, 1901</synonym>
	<dcterms:created>1998-04-30</dcterms:created>
	<dcterms:modified>2014-05-15</dcterms:modified>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=593&amp;speccode=48090'>Berg, L.S.0 Freshwater fishes of the U.S.S.R. and adjacent countries. volume 1, 4th edition. Israel Program for Scientific Translations Ltd, Jerusalem. (Russian version published 1948). (Ref. 593)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=12801&amp;speccode=48090'>Svetovidov, A.N.0 Clupeidae. p. 99-109. In J.C. Hureau and Th. Monod (eds.) Check-list of the fishes of the north-eastern Atlantic and of the Mediterranean (CLOFNAM). UNESCO, Paris. Vol. 1. (Ref. 12801)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=13696&amp;speccode=48090'>Kottelat, M.0 European freshwater fishes. An heuristic checklist of the freshwater fishes of Europe (exclusive of former USSR), with an introduction for non-systematists and comments on nomenclature and conservation. Biologia, Bratislava, 52/Suppl. 5:1-271. (Ref. 13696)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=26281&amp;speccode=48090'>Balon, E.K.0 Reproductive guilds of fishes: a proposal and definition. J. Fish. Res. Board Can. 32(6):821-864. (Ref. 26281)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=26282&amp;speccode=48090'>Eschmeyer, W.N. (ed.)0 Catalog of fishes. Special Publication, California Academy of Sciences, San Francisco. 3 vols. 2905 p. (Ref. 26282)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=26334&amp;speccode=48090'>Reshetnikov, Y.S., N.G. Bogutskaya, E.D. Vasil'eva, E.A. Dorofeeva, A.M. Naseka, O.A. Popova, K.A. Savvaitova, V.G. Sideleva and L.I. Sokolov0 An annotated check-list of the freshwater fishes of Russia. J. Ichthyol. 37(9):687-736. (Ref. 26334)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=37297&amp;speccode=48090'>Mikhov, S.0 Checklist of fishes of Bulgaria. FishBase checklist modified by Stoyan Mikhov. (Ref. 37297)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=38732&amp;speccode=48090'>Anonymous0 Fish collection database of the National Museum of Natural History (Smithsonian Institution). Smithsonian Institution-Division of Fishes. (Ref. 38732)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=39150&amp;speccode=48090'>Banarescu, P.0 Pisces - Osteichthyes (pesti ganoizi si osisi). Editura Academiei Republicii Populare Romîne, Bucuresti. 962 p. (Ref. 39150)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=41476&amp;speccode=48090'>Deval, M.C.0 Common names of marine fishes of Turkey. Unpublished report. (Ref. 41476)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=45043&amp;speccode=48090'>Bogutskaya, N.G.0 Common names of Russian freshwater fishes. Spreadsheet with cyrillic names made available for inclusion in FishBase. Unpublished. (Ref. 45043)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=47438&amp;speccode=48090'>Anonymous0 Fish collection of the Royal Ontario Museum. Royal Ontario Museum. (Ref. 47438)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=51054&amp;speccode=48090'>Hanel, L. and J. Novák0 České názvy zivočichů V. Ryby a rybovití obratlovci (Pisces) II., nozdratí (Sarcopterygii), paprskoploutví (Actinopterygii) [chrupavčití (Chondrostei), kostnatí (Neopterygii): kostlíni (Semionotiformes) - bezostní (Clupeiformes)]. Národní muzeum (zoologické odděleí), Praha. (Ref. 51054)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=51243&amp;speccode=48090'>Riede, K.0 Global register of migratory species - from global to regional scales. Final Report of the R&amp;D-Projekt 808 05 081. Federal Agency for Nature Conservation, Bonn, Germany. 329 p. (Ref. 51243)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=56563&amp;speccode=48090'>Moskvin, B.S.0 Materials on the feeding of shads in the northeastern part of the Black Sea. Trudy Novorossiiskoi Biologicheskoi Stantsii 2(3):259-272. (Ref. 56563)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=57855&amp;speccode=48090'>Bilecenoğlu, M., Taskavak E., S. Mater and M. Kaya0 Checklist of the marine fishes of Turkey. Zootaxa 113(1):1-194. (Ref. 57855)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=58293&amp;speccode=48090'>Bogutskaya, N.G. and A.M. Naseka0 REGIONAL CHECK-LISTS: Don River Drainage Area. In Website and Database: "Freshwater Fishes of Russia": A Source of Information on the Current State of the Fauna. Zoological Institute RAS. (Ref. 58293)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=58296&amp;speccode=48090'>Bogutskaya, N.G. and A.M. Naseka0 REGIONAL CHECK-LISTS: Kuban River Drainage Area. In Website and Database: "Freshwater Fishes of Russia": A Source of Information on the Current State of the Fauna. Zoological Institute RAS. (Ref. 58296)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=58342&amp;speccode=48090'>Fricke, R., M. Bilecenoglu and H.M. Sari0 Annotated checklist of fish and lamprey species (Gnathostoma and Petromyzontomorphi) of Turkey, including a Red List of threatened and declining species. Stuttgarter Beitr. Naturk. Sea A (706):1-172. (Ref. 58342)</reference>
	<reference isbn='978-2-8399-0298-4' url='http://www.fishbase.org/references/FBRefSummary.php?id=59043&amp;speccode=48090'>Kottelat, M. and J. Freyhof0 Handbook of European freshwater fishes. Publications Kottelat, Cornol and Freyhof, Berlin. 646 pp. (Ref. 59043)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=59105&amp;speccode=48090'>Tarkan, A.S., O. Gaygusuz, H. Acipinar, C. Gürsoy and M. Ozulug0 Length-weight relationship of fishes from the Marmara region (NW-Turkey). J. Appl. Ichthyol. 22:271-273. (Ref. 59105)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=88083&amp;speccode=48090'>Bat, L., Y. Erdem, S. Ustaoğlu, Ö Yardım and H. Hüseyin Satılmış0 A study on the fishes of the Central Black Sea coast of Turkey. J. Black Sea/Mediterranean Environment 11:291-296. (Ref. 88083)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=96734&amp;speccode=48090'>Ergudren, E., F. Turan and C. Turan0 Length-weight and length-length relationships for four shad species along the western Black Sea coast of Turkey. J. Appl. Ichthyol. 27:942-944. (Ref. 96734)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=105471&amp;speccode=48090'>eElurikkus0 eElurikkus koondab ühte portaali Eesti eluslooduse andmed. http://elurikkus.ut.ee/ (Ref. 105471)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=106151&amp;speccode=48090'>Çiçek, E., R. Fricke and S.S. Birecikligil0 Freshwater fishes of Turkey: a revised and updated annotated checklist. Biharean Biologist 9(2):141-157. (Ref. 106151)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=126062&amp;speccode=48090'>Dyldin, Y.V., A.M. Orlov, L. Hanel, V. Romanov, R. Fricke and E.D. Vasil'eva0 Ichthyofauna of fresh and brackish waters of Russia and adjacent areas: annotated list with taxonomic comments: 1. Families Petromyzontidae-Pristigasteridae. J. Ichthyol. 2022:1-31. (Ref. 126062)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=128151&amp;speccode=48090'>Abliazov, E.R., A.R. Boltachev, E.P. Karpova, A.N. Pashkov and O.N. Danilyuk0 Ichthyofauna of the Black Sea coastal zone in the Laspi Bay area (Crimea). Mar. Biol. J. 6(2):3-17. (Ref. 128151)</reference>
	<reference isbn='2350-4757' url='http://www.fishbase.org/references/FBRefSummary.php?id=128520&amp;speccode=48090'>Marčeta, B. and K. Pliberšek (eds)0 BiosWeb. Biological database of the Fisheries Research Institute of Slovenia. Ljubljana, Fisheries Research Institute of Slovenia, 2014. (www.biosweb.org. updated 13/04/2023). (Ref. 128520)</reference>
	<reference isbn='1849954984' url='http://www.fishbase.org/references/FBRefSummary.php?id=128868&amp;speccode=48090'>Beolens, B., M. Grayson and M. Watkins0 Eponym dictionary of Fishes. 	Dunbeath, Caithness (Scotland, UK): Whittles Publishing Limited, 1300 p. (Ref. 128868)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=130435&amp;speccode=48090'>IUCN0 The IUCN Red List of Threatened Species. 1964 - onwards. https://www.iucnredlist.org/ (Ref. 130435)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=138126&amp;speccode=48090'>Lassalle, G., M. Béguer, L. Baulaton and E. Rochard0 Learning from the past to predict the future: Responses of European diadromous fish to climate change. Amer. Fish. Soc. Symp. 69:175-193. (Ref. 138126)</reference><dataObject>
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	<dcterms:created>1998-04-30 00:00:00</dcterms:created>
	<dcterms:modified>2025-10-21 00:00:00</dcterms:modified>
	<dc:title>Biology</dc:title>
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	<dc:description>This species migrates from sea to mouth and lower reaches of large rivers (at 1-2 years) and spawns in fresh or slightly brackish water, usually close to the shore, upper 2-4 m, in almost still water bodies as flood plains or lakes.  Many individuals spawn 2-4 seasons and spawners appear along the coast in late January to March, enter rivers when temperature reaches about 10°C in late April to May, and spawns at about 15°C or beyond in May to June.  Eggs are bathypelagic or sink to bottom.  Spent fish migrate to estuarine and coastal lagoons or to sea near river mouths to feed and in autumn, move to sea near river mouths to overwinter.  Juveniles migrate to sea or estuaries during first summer and remain there until maturity.  At sea, feeds on a wide variety of zooplankton (crustaceans), insect larvae and small fish.  Populations seem to have stabilized at moderate level after most have declined during the first decades of 20th century due to water pollution.  In northern Black Sea, immediate impact is foreseen due to the decline in habitat quality in suitable estuarine ecosystems.  On the other hand, the species is still abundant in Azov Sea basin (Ref. 59043).</dc:description>
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	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=593&amp;speccode=48090'>Berg, L.S. 1962 Freshwater fishes of the U.S.S.R. and adjacent countries. volume 1, 4th edition. Israel Program for Scientific Translations Ltd, Jerusalem. (Russian version published 1948). (Ref. 593)</reference></dataObject><dataObject>
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	<agent homepage='' logoURL='' role='recorder'>Estelita Emily Capuli</agent>
	<dcterms:created>2010-07-13 00:00:00</dcterms:created>
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	<dc:description>Distinguished from its congeners entering freshwater in Black Sea basins by having 66-96 gill rakers, longer than branchial filaments and poorly-developed teeth on palatine and vomer (Ref. 59043).</dc:description>
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	<reference isbn='978-2-8399-0298-4' url='http://www.fishbase.org/references/FBRefSummary.php?id=59043&amp;speccode=48090'>Kottelat, M. and J. Freyhof 2007 Handbook of European freshwater fishes. Publications Kottelat, Cornol and Freyhof, Berlin. 646 pp. (Ref. 59043)</reference></dataObject><dataObject>
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	<dc:description>Eurasia:  Black Sea and Sea of Azov from where adults ascend rivers, migrating a short distance upstream to spawn.</dc:description>
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	<reference isbn='978-2-8399-0298-4' url='http://www.fishbase.org/references/FBRefSummary.php?id=59043&amp;speccode=48090'>Kottelat, M. and J. Freyhof 2007 Handbook of European freshwater fishes. Publications Kottelat, Cornol and Freyhof, Berlin. 646 pp. (Ref. 59043)</reference></dataObject><dataObject>
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	<dc:description>Anadromous. Fish that ascend rivers to spawn, as salmon and hilsa do. Sub-division of diadromous.    Migrations should be cyclical and predictable and cover more than 100 km.</dc:description>
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	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=138126&amp;speccode=48090'>Lassalle, G., M. Béguer, L. Baulaton and E. Rochard 2009 Learning from the past to predict the future: Responses of European diadromous fish to climate change. Amer. Fish. Soc. Symp. 69:175-193. (Ref. 138126)</reference></dataObject><dataObject>
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	<dc:description>Juveniles migrate to sea or estuaries during their first summer, remaining there until they mature.  After 1-2 years, they migrate upriver to breed. Spawners appear along the coast in late January-March, enter rivers when temperature reaches about 10°C, in late April and May.  Spawning occurs from May to June on or above 15°C.  Spent individuals migrate to estuaries and coastal lagoons or to sea near river mouths to feed.  In autumn, they move to the sea to overwinter (Ref. 59043).</dc:description>
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	<dc:description>30.0 cm TL (male/unsexed; (Ref. 96734)); max. published weight: 59.0 g (Ref. 593)</dc:description>
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	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=96734&amp;speccode=48090'>Ergudren, E., F. Turan and C. Turan 2011 Length-weight and length-length relationships for four shad species along the western Black Sea coast of Turkey. J. Appl. Ichthyol. 27:942-944. (Ref. 96734)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=593&amp;speccode=48090'>Berg, L.S. 1962 Freshwater fishes of the U.S.S.R. and adjacent countries. volume 1, 4th edition. Israel Program for Scientific Translations Ltd, Jerusalem. (Russian version published 1948). (Ref. 593)</reference></dataObject><dataObject>
	<dc:identifier>FB-Threats-2</dc:identifier>
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	<agent homepage='' logoURL='' role='recorder'>Belen Acosta</agent>
	<dcterms:created>1990-10-18 00:00:00</dcterms:created>
	<dcterms:modified>2024-03-07 00:00:00</dcterms:modified>
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	<license>http://creativecommons.org/licenses/by-nc/3.0/</license>
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	<dcterms:rightsHolder> FAO</dcterms:rightsHolder>
	<dcterms:bibliographicCitation></dcterms:bibliographicCitation>
	<audience>General public</audience>
	<dc:source>http://www.fishbase.org/summary/SpeciesSummary.php?id=2</dc:source>
	<subject></subject>
	<dc:description>Least Concern (LC) , IUCN Grouper and Wrasse Specialist Group</dc:description>
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	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=57073'>IUCN 2006 2006 IUCN red list of threatened species. www.iucnredlist.org. Downloaded July 2006.</reference></dataObject><dataObject>
	<dc:identifier>FB-Habitat-48090</dc:identifier>
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	<agent homepage='' logoURL='' role='recorder'>Pascualita Sa-a</agent>
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	<dcterms:modified>2025-10-21 00:00:00</dcterms:modified>
	<dc:title>Environment</dc:title>
	<dc:language></dc:language>
	<license>http://creativecommons.org/licenses/by-nc/3.0/</license>
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	<dcterms:rightsHolder>WorldFish Center - FishBase</dcterms:rightsHolder>
	<dcterms:bibliographicCitation></dcterms:bibliographicCitation>
	<audience>General public</audience>
	<dc:source>http://www.fishbase.org/summary/SpeciesSummary.php?id=48090</dc:source>
	<subject>http://rs.tdwg.org/ontology/voc/SPMInfoItems#Distribution</subject>
	<dc:description>pelagic-neritic; anadromous (Ref. 138126); freshwater; brackish; marine; depth range 50 - 70 m (Ref. 59043)</dc:description>
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	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=138126&amp;speccode=48090'>Lassalle, G., M. Béguer, L. Baulaton and E. Rochard 2009 Learning from the past to predict the future: Responses of European diadromous fish to climate change. Amer. Fish. Soc. Symp. 69:175-193. (Ref. 138126)</reference>
	<reference isbn='978-2-8399-0298-4' url='http://www.fishbase.org/references/FBRefSummary.php?id=59043&amp;speccode=48090'>Kottelat, M. and J. Freyhof 2007 Handbook of European freshwater fishes. Publications Kottelat, Cornol and Freyhof, Berlin. 646 pp. (Ref. 59043)</reference></dataObject>
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