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<taxon>
	<dc:identifier>FB-4785</dc:identifier>
	<dc:source>http://www.fishbase.org/summary/SpeciesSummary.php?id=4785</dc:source>
	<dwc:Kingdom>Animalia</dwc:Kingdom>
	<dwc:Phylum>Chordata</dwc:Phylum>
	<dwc:Class>Teleostei</dwc:Class>
	<dwc:Order>Cypriniformes</dwc:Order>
	<dwc:Family>Leuciscidae</dwc:Family>
	<dwc:Genus>Pimephales</dwc:Genus>
	<dwc:ScientificName>Pimephales promelas Rafinesque, 1820</dwc:ScientificName>
	<commonName xml:lang='cs'>Jeleček velkohlavý</commonName>
	<commonName xml:lang='da'>Tykhovedet elritse</commonName>
	<commonName xml:lang='da'>Tyrehoved</commonName>
	<commonName xml:lang='de'>Fettköpfige Elritze</commonName>
	<commonName xml:lang='en'>Black-head minnow</commonName>
	<commonName xml:lang='en'>Fathead minnow</commonName>
	<commonName xml:lang='en'>Rosey reds</commonName>
	<commonName xml:lang='et'>Tüse tömpnina</commonName>
	<commonName xml:lang='fi'>Paksupäämutu</commonName>
	<commonName xml:lang='fr'>Tête de boule</commonName>
	<commonName xml:lang='fr'>Tête-de-boule</commonName>
	<commonName xml:lang='sv'>Knölskallelöja</commonName>
	<commonName xml:lang='es'>Carpita cabezona</commonName>
	<commonName xml:lang='es'>Mino cabezón</commonName>
	<commonName xml:lang=''>Tolstoglavi pisanec</commonName>
	<commonName xml:lang=''>胖头鱥</commonName>
	<commonName xml:lang=''>胖頭鱥</commonName>
	<synonym relationship='synonym'>Cliola smithii Evermann &amp; Cox, 1896</synonym>
	<synonym relationship='synonym'>Colistus parietalis Cope, 1871</synonym>
	<synonym relationship='synonym'>Hyborhynchus confertus Girard, 1856</synonym>
	<synonym relationship='synonym'>Hyborhynchus nigellus Cope, 1875</synonym>
	<synonym relationship='synonym'>Hyborhynchus puniceus Girard, 1856</synonym>
	<synonym relationship='synonym'>Pimephales agassizii Cope, 1867</synonym>
	<synonym relationship='synonym'>Pimephales anuli Kendall, 1903</synonym>
	<synonym relationship='synonym'>Pimephales fasciatus Girard, 1856</synonym>
	<synonym relationship='synonym'>Pimephales maculosus Girard, 1856</synonym>
	<synonym relationship='synonym'>Pimephales milesii Cope, 1865</synonym>
	<synonym relationship='synonym'>Pimephales promelas harveyensis Hubbs &amp; Lagler, 1949</synonym>
	<synonym relationship='synonym'>Pimephales promelas promelas Rafinesque, 1820</synonym>
	<synonym relationship='synonym'>Pimephalus promelas Rafinesque, 1820</synonym>
	<synonym relationship='synonym'>Plargyrus melanocephalus Abbott, 1860</synonym>
	<dcterms:created>1991-11-28</dcterms:created>
	<dcterms:modified>2024-01-15</dcterms:modified>
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	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=77059&amp;speccode=4785'>Brown, G.E., D.P. Chivers and R.J.F. Smith0 Fathead Minnows avoid conspecific and heterospecific alarm pheromones in the feces of northern pike. J. Fish Biol. 47(3):387-393. (Ref. 77059)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=77062&amp;speccode=4785'>Kusch, R.C., R.S. Mirza and D.P. Chivers0 Making sense of predator scents: investigating the sophistication of predator assessment abilities of fathead minnows. Behavioral Ecology and Sociobiology 55:551-555. (Ref. 77062)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=77066&amp;speccode=4785'>Chivers, D.P. and R.J.F. Smith0 The role of experience and chemical alarm signalling in predator recognition by fathead minnows, &lt;i&gt;Pimephales promelas&lt;/i&gt;. J. Fish Biol. 44(2):273-285. (Ref. 77066)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=77067&amp;speccode=4785'>Chivers, D.P. and R.J.F. Smith0 Free-living fathead minnows rapidly learn to recognize pike as predators. J. Fish Biol. 46(6):949-954. (Ref. 77067)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=77068&amp;speccode=4785'>Chivers, D.P. and R.J.F. Smith0 Fathead minnows (&lt;i&gt;Pimephales promelas&lt;/i&gt;) learn to recognize chemical stimuli from high-risk habitats by the presence of alarm substance. Behavioral Ecology 6:155-158. (Ref. 77068)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=78789&amp;speccode=4785'>Missouri Department of Conservation0 Fish of Missouri. Missouri Department of Conservation. (Ref. 78789)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=78807&amp;speccode=4785'>Lieb, C.S.0 Annotated checklist of the fishes of the Rio Grande drainage, Dona Ana, El Paso, and Hudspeth counties. Centennial Museum, University of Texas at El Paso. (Ref. 78807)</reference>
	<reference isbn='0-88864-236-9' url='http://www.fishbase.org/references/FBRefSummary.php?id=78981&amp;speccode=4785'>Nelson, J.S. and M.J. Paetz0 The fishes of Alberta. 2nd edit. Edmonton (AB, Canada): The University of Alberta Press. (Ref. 78981)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=78982&amp;speccode=4785'>Rempel, L.L. and D.G. Smith0 Postglacial fish dispersal from the Mississippi refuge to the Mackenzie River basin. Can. J. Fish. Aquat. Sci. 55:893-899. (Ref. 78982)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=79012&amp;speccode=4785'>Hassan-Williams, C., T.H. Bonner and C. Thomas (photogr.)0 Texas freshwater fishes. Texas State University-San Marcos: Biology Department/ Aquatic Station. (Ref. 79012)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=79522&amp;speccode=4785'>Verreycken, H., D. Anseeuw, G. Van Thuyne, P. Quataert and C. Belpaire0 The non-indigenous freshwater fishes of Flanders (Belgium): review, status and trends over the last decade. J. Fish Biol. 71(Suppl.D):160-172. (Ref. 79522)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=83882&amp;speccode=4785'>FAO-FIES0 Aquatic Sciences and Fisheries Information System (ASFIS) species list. Retrieved from http://www.fao.org/fishery/collection/asfis/en, March 2010. (Ref. 83882)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=86798&amp;speccode=4785'>Page, L.M. and B.M. Burr0 A field guide to freshwater fishes of North America north of Mexico. Boston : Houghton Mifflin Harcourt, 663p. (Ref. 86798)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=88166&amp;speccode=4785'>Verreycken, H., G. Van Thuyne and C. Belpaire0 Length-weight relationships of 40 freshwater fish species from two decades of monitoring in Flanders (Belgium). J. Appl. Ichthyol. 2011:1-5. (Ref. 88166)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=90062&amp;speccode=4785'>FAO-FIES0 Aquatic Sciences and Fisheries Information System (ASFIS) species list. Retrieved from http://www.fao.org/fishery/collection/asfis/en, March 2012. (Ref. 90062)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=95632&amp;speccode=4785'>FAO-FIES0 Aquatic Sciences and Fisheries Information System (ASFIS) species list. Retrieved from http://www.fao.org/fishery/collection/asfis/en, April 2014. (Ref. 95632)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=101110&amp;speccode=4785'>FAO-FIES0 Aquatic Sciences and Fisheries Information System (ASFIS) species list. Retrieved from http://www.fao.org/fishery/collection/asfis/en, [accessed 13/04/2015]. (Ref. 101110)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=105471&amp;speccode=4785'>eElurikkus0 eElurikkus koondab ühte portaali Eesti eluslooduse andmed. http://elurikkus.ut.ee/ (Ref. 105471)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=112251&amp;speccode=4785'>Taylor, E.B. and K. Tan0 Guide to the freshwater fishes of British Columbia. 1st edit. Vancouver (BC, Canada): Beaty Biodiversity Museum, 18 p. (Ref. 112251)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=115257&amp;speccode=4785'>FAO-FIES0 Aquatic Sciences and Fisheries Information System (ASFIS) species list. Retrieved from http://www.fao.org/fishery/collection/asfis/en (accessed 08/06/2017). (Ref. 115257)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=118272&amp;speccode=4785'>FAO-FIES0 Aquatic Sciences and Fisheries Information System (ASFIS) species list. Retrieved from htttps://www.fao.org/fishery/collection/asfis/en [accessed May, 2018] (Ref. 118272)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=119194&amp;speccode=4785'>Tonn, W., H. Swanson, C. Paszkowski, J. Hanisch and L. Chavarie0 Northern North America. p. 85-100. In J.F. Craig (ed.).  Freshwater Fisheries Ecology. John Wiley &amp; Sons Ltd. 899p. (Ref. 119194)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=126106&amp;speccode=4785'>Dean, E.M., A.R. Cooper, L. Wang, W. Daniel, S. David, C. Ernzen, K.B. Gido, E Hale, T.J. Haxton, W. Kelso, N. Leonard, C. Lido, J. Margraf, M. Porter, C. Pennock, D. Propst, J. Ross, M.D. Staudinger, G. Whelan and D.M. Infante0 The North American Freshwater Migratory Fish Database (NAFMFD): Characterizing the migratory life histories of freshwater fishes of Canada, the United States of Mexico. J. Biogeography 49:1193-1203. (Ref. 126106)</reference>
	<reference isbn='2350-4757' url='http://www.fishbase.org/references/FBRefSummary.php?id=128520&amp;speccode=4785'>Marčeta, B. and K. Pliberšek (eds)0 BiosWeb. Biological database of the Fisheries Research Institute of Slovenia. Ljubljana, Fisheries Research Institute of Slovenia, 2014. (www.biosweb.org. updated 13/04/2023). (Ref. 128520)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=132215&amp;speccode=4785'>WDNR, Wisconsin Department of Natural Resources0 Wisconsin Fish Species List.. World Wide Web electronic publication, Accessed 05 Oct. 2024. Madison (USA, WI):  Wisconsin Department of Natural Resources, 6 p. https://dnr.wisconsin.gov/sites/default/files/topic/Fishing/Species_wifish.pdf (Ref. 132215)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=137577&amp;speccode=4785'>Price, S.A., S.T. Friedman, K.A. Corn, O. Larouche, K. Brockelsby, A.J. Lee, M. Nagaraj, N.G. Bertrand, M. Danao, M.C. Coyne, J.R. Estrada, R. Friedman, E. Hoeft, M. Iwan, D. Gross, J.H. Kao, B. Landry, M.J. Linares, C. McGlinn and …0 FishShapes v1: Functionally relevant measurements of teleost shape and size on three dimensions. Ecology 103(e3829):1-2. (Ref. 137577)</reference><dataObject>
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	<dc:description>Adults inhabit muddy pools of headwaters, creeks, small rivers, and ponds (Ref. 5723, 86798).  Also found in lakes (Ref. 10294).  Can tolerate conditions (e.g., turbid, hot, poorly oxygenated, intermittent streams) unsuitable for most fishes (Ref. 5723, 86798).  They feed on detritus and algae (Ref. 10294).  Spawning occurs in still-water habitats along shores (Ref. 59043).  Introductions consequently caused the spread of the enteric red-mouth disease throughout northern Europe which infected wild and cultured trouts and eels (Ref. 1739).  Maintained a relatively high metabolic rate and level of activity under hypoxic conditions (Ref. 77050).  Individuals  that survived the hypoxic conditions during winter had rapid growth rates after ice-off (Ref. 77048).</dc:description>
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	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=86798&amp;speccode=4785'>Page, L.M. and B.M. Burr 2011 A field guide to freshwater fishes of North America north of Mexico. Boston : Houghton Mifflin Harcourt, 663p. (Ref. 86798)</reference></dataObject><dataObject>
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	<dc:description>The only species of genus introduced to Europe which differs from other cyprinid species in Europe by the following characters:  prominent pad of spongy tissue on nape in males; dorsal fin origin about above pelvic fin origin; anal fin with 7½ branched rays; second simple dorsal ray spinous, about 50% length of the third one; lateral line incomplete, usually not reaching dorsal origin; weak midlateral stripe; 41-54 + 2 scales in midlateral row; and mouth terminal (Ref. 59043).</dc:description>
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	<reference isbn='2-85653653261' url='http://www.fishbase.org/references/FBRefSummary.php?id=40476&amp;speccode=4785'>Keith, P. and J. Allardi (coords.) 2001 Atlas des poissons d'eau douce de France. Muséum national d’Histoire naturelle, Paris. Patrimoines naturels, 47:1-387. (Ref. 40476)</reference></dataObject><dataObject>
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	<dc:description>Dorsal spines (total): 0; Dorsal soft rays (total): 8;  Analspines: 0; Analsoft rays: 7</dc:description>
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	<reference isbn='2-85653653261' url='http://www.fishbase.org/references/FBRefSummary.php?id=40476&amp;speccode=4785'>Keith, P. and J. Allardi (coords.) 2001 Atlas des poissons d'eau douce de France. Muséum national d’Histoire naturelle, Paris. Patrimoines naturels, 47:1-387. (Ref. 40476)</reference></dataObject><dataObject>
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	<dc:description>North and Central America:  Over much of North America from Quebec to Northwest Territories and British Columbia in Canada and south to Alabama, Texas in USA, and Mexico.  Widely introduced, including in Colorado River drainage in Arizona and New Mexico, USA.  Not present on Atlantic and Gulf slopes between Potomac River in Virginia and Trinity River in Texas.  Several countries report adverse ecological impact after introduction.</dc:description>
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	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=86798&amp;speccode=4785'>Page, L.M. and B.M. Burr 2011 A field guide to freshwater fishes of North America north of Mexico. Boston : Houghton Mifflin Harcourt, 663p. (Ref. 86798)</reference></dataObject><dataObject>
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	<dc:title>Importance</dc:title>
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	<subject>http://rs.tdwg.org/ontology/voc/SPMInfoItems#Size</subject>
	<dc:description>fisheries: commercial; aquarium: commercial; bait: usually</dc:description>
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	<agent homepage='' logoURL='' role='recorder'>Armi G. Torres</agent>
	<dcterms:created>1991-11-28 00:00:00</dcterms:created>
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	<dc:source>http://www.fishbase.org//Reproduction/FishReproSummary.php?ID=4785&amp;StockCode=5009</dc:source>
	<subject>http://rs.tdwg.org/ontology/voc/SPMInfoItems#Threats</subject>
	<dc:description>"Males defend territories and eggs, and clean plants, stone or other materials by rubbing them with the spongy nape pad and dorsal fin, which contain mucus-secreting cells and taste buds.  The mucus may have fungicidal properties" (Ref. 59043).</dc:description>
	<mediaURL></mediaURL>
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	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=26281&amp;speccode=4785'>Balon, E.K. 1975 Reproductive guilds of fishes: a proposal and definition. J. Fish. Res. Board Can. 32(6):821-864. (Ref. 26281)</reference></dataObject><dataObject>
	<dc:identifier>FB-TrophicStrategy-4785</dc:identifier>
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	<agent homepage='' logoURL='' role='recorder'>Pascualita Sa-a</agent>
	<dcterms:created>1997-04-08 00:00:00</dcterms:created>
	<dcterms:modified>2008-07-17 00:00:00</dcterms:modified>
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	<dcterms:bibliographicCitation></dcterms:bibliographicCitation>
	<audience>General public</audience>
	<dc:source>promelas</dc:source>
	<subject>http://rs.tdwg.org/ontology/voc/SPMInfoItems#Habitat</subject>
	<dc:description>Inhabits muddy pools of headwaters, creeks and small rivers.  Also found in ponds.  Tolerates unsuitable conditions (e.g., turbid, hot, poorly oxygenated, intermittent streams).   Interannual population density varied dramatically in response to the severity of the preceding winter for species inhabiting boreal lakes in Alberta, Canada (Ref. 77054).  Populations from two shallow lakes repeatedly winterkilled, 47-94% post winterkill declines in density, increased growth rates of the surviving fish (Ref. 77054).  Maintained a relatively high metabolic rate and level of activity under hypoxic conditions (Ref. 77050).  Individuals  that survived the hypoxic conditions during winter had rapid growth rates after ice-off (Ref. 77048). Responds to visual and chemical cues of northern pike, Esox lucius.  Species found in naturally predator-free waters learned to avoid the 'scent' of introduced pike faster than the sight of them (Ref. 77049).  Anti-predator responses by fathead minnows increased with higher concentrations of chemical "odor" emitted from northern pike.  More frightened by the cues of smaller-sized pike, which may pose a greater predation threat than larger pike (Ref. 77062).</dc:description>
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	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=1998&amp;speccode=4785'>Scott, W.B. and E.J. Crossman 1973 Freshwater fishes of Canada. Bull. Fish. Res. Board Can. 184: xi+1-966. (Ref. 1998)</reference></dataObject><dataObject>
	<dc:identifier>FB-Size-4785</dc:identifier>
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	<agent homepage='' logoURL='' role='recorder'>Susan M. Luna</agent>
	<dcterms:created>1991-11-28 00:00:00</dcterms:created>
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	<dc:title>Max. size</dc:title>
	<dc:language></dc:language>
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	<audience>General public</audience>
	<dc:source>http://www.fishbase.org/summary/SpeciesSummary.php?id=4785</dc:source>
	<subject>http://rs.tdwg.org/ontology/voc/SPMInfoItems#DiagnosticDescription</subject>
	<dc:description>10.1 cm TL (male/unsexed; (Ref. 10294)); max. reported age: 5 years (Ref. 58435)</dc:description>
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	<reference isbn='0-87049-711-1' url='http://www.fishbase.org/references/FBRefSummary.php?id=10294&amp;speccode=4785'>Etnier, D.A. and W.C. Starnes 1993 The fishes of Tennessee. The University of Tennessee Press, Knoxville, Tennessee, USA. (pls. check date). (Ref. 10294)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=58435&amp;speccode=4785'>Danylchuk, A.J. and W.M. Tonn 2006 Natural disturbance and life history: consequences of winterkill on fathead minnow on boreal lakes. J. Fish Biol. 68(3):681-694. (Ref. 58435)</reference></dataObject><dataObject>
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	<agent homepage='' logoURL='' role='recorder'>Belen Acosta</agent>
	<dcterms:created>1990-10-18 00:00:00</dcterms:created>
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	<dc:source>http://www.fishbase.org/summary/SpeciesSummary.php?id=2</dc:source>
	<dc:description>Least Concern (LC) , IUCN Grouper and Wrasse Specialist Group</dc:description>
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	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=57073'>IUCN 2006 2006 IUCN red list of threatened species. www.iucnredlist.org. Downloaded July 2006.</reference></dataObject><dataObject>
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	<agent homepage='' logoURL='' role='recorder'>Allan Palacio</agent>
	<dcterms:created>2004-03-17 00:00:00</dcterms:created>
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	<audience>General public</audience>
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	<subject>http://rs.tdwg.org/ontology/voc/SPMInfoItems#Distribution</subject>
	<dc:description>Enteric Redmouth Disease. Bacterial diseases</dc:description>
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	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=48849&amp;speccode=4785'>Horne, M.T. and A.C. Barnes 1999 Enteric redmouth disease (&lt;i&gt;Yersinia ruckeri&lt;/i&gt;). p.455-477. In P.T.K. Woo and D.W. Bruno (eds.) Fish Diseases and Disorders, Vol. 3: Viral, Bacterial and Fungal Infections. CAB Int'l. (Ref. 48849)</reference></dataObject><dataObject>
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	<agent homepage='' logoURL='' role='recorder'>Susan M. Luna</agent>
	<dcterms:created>1991-11-28 00:00:00</dcterms:created>
	<dcterms:modified>2025-12-04 00:00:00</dcterms:modified>
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	<dc:source>http://www.fishbase.org/summary/SpeciesSummary.php?id=4785</dc:source>
	<dc:description>demersal; non-migratory (Ref. 126106); freshwater</dc:description>
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	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=126106&amp;speccode=4785'>Dean, E.M., A.R. Cooper, L. Wang, W. Daniel, S. David, C. Ernzen, K.B. Gido, E Hale, T.J. Haxton, W. Kelso, N. Leonard, C. Lido, J. Margraf, M. Porter, C. Pennock, D. Propst, J. Ross, M.D. Staudinger, G. Whelan and D.M. Infante 2022 The North American Freshwater Migratory Fish Database (NAFMFD): Characterizing the migratory life histories of freshwater fishes of Canada, the United States of Mexico. J. Biogeography 49:1193-1203. (Ref. 126106)</reference></dataObject>
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