FB-4779 http://www.fishbase.org/summary/SpeciesSummary.php?id=4779 Animalia Chordata Teleostei Anabantiformes Anabantidae Ctenopoma Ctenopoma nigropannosum Reichenow, 1875 Toplettet buskfisk Zweifleck-Buschfisch Twospot climbing perch Twospot ctenopoma Täpläkiipijäkala Torpedokiipijäkala Ekenge Gehenge Ikekenge 双点非洲攀鲈 雙點非洲攀鱸 Anabas nigropannosa (Reichow, 1875) Anabas nigropannosus (Reicherow, 1875) 1991-11-28 2017-10-16 Breder, C.M. and D.E. Rosen0 Modes of reproduction in fishes. T.F.H. Publications, Neptune City, New Jersey. 941 p. (Ref. 205) Welcomme, R.L.0 International introductions of inland aquatic species. FAO Fish. Tech. Pap. 294. 318 p. (Ref. 1739) Boulenger, G.A.0 Catalogue of the fresh-water fishes of Africa in the British Museum (Natural History). Volume IV. London. 392 p. (Ref. 1880) Courtenay, W.R. Jr. and C.R. Robins0 Exotic aquatic organisms in Florida with emphasis on fishes: a review and recommendations. Trans. Am. Fish. Soc. 102(1):1-12. (Ref. 1987) Baensch, H.A. and R. Riehl0 Aquarien atlas. Band 2. Mergus, Verlag für Natur-und Heimtierkunde GmbH, Melle, Germany. 1216 p. (Ref. 2060) Robins, C.R., R.M. Bailey, C.E. Bond, J.R. Brooker, E.A. Lachner, R.N. Lea and W.B. Scott0 Common and scientific names of fishes from the United States and Canada. Am. Fish. Soc. Spec. Publ. (20):183 p. (Ref. 3814) Hureau, J.-C.0 La base de données GICIM: Gestion informatisée des collections ichthyologiques du Muséum. p. 225-227. In Atlas Préliminaire des Poissons d'Eau Douce de France. Conseil Supérieur de la Pêche, Ministère de l'Environnement, CEMAGREF et Muséum national d'Histoire naturelle, Paris. (Ref. 4517) Robins, C.R., R.M. Bailey, C.E. Bond, J.R. Brooker, E.A. Lachner, R.N. Lea and W.B. Scott0 World fishes important to North Americans. Exclusive of species from the continental waters of the United States and Canada. Am. Fish. Soc. Spec. Publ. (21):243 p. (Ref. 4537) Gosse, J.-P.0 Anabantidae. p. 402-414. In J. Daget, J.-P. Gosse and D.F.E. Thys van den Audenaerde (eds.) Check-list of the freshwater fishes of Africa (CLOFFA). ISNB, Brussels; MRAC, Tervuren; and ORSTOM, Paris. Vol. 2. (Ref. 6803) Courtenay, W.R. Jr., H.F. Sahlman, W.W. Miley II and D.J. Herrema0 Exotic fishes in fresh and brackish waters of Florida. Biological Conservation. 6(4): 292-304. (Ref. 10757) Anonymous0 Fish registrations within the museum database of the Vertebrate Section of the Royal Museum for Central Africa. MRAC, Tervuren, Belgium. (Ref. 12818) Wu, H.L., K.-T. Shao and C.F. Lai (eds.)0 Latin-Chinese dictionary of fishes names. The Sueichan Press, Taiwan. 1028 p. (Ref. 31517) Anonymous0 Fish collection database of the Natural History Museum, London (formerly British Museum of Natural History (BMNH)). Natural History Museum, London (formerly British Museum of Natural History (BMNH)). (Ref. 31982) Anonymous0 The icthyological collection of the Zoological Museum Hamburg (ZMH). Division of Ichthyology and Herpetology, Zoological Museum Hamburg (ZMH). (Ref. 35508) Anonymous0 Fish collection database of the American Museum of Natural History. American Museum of Natural History, Central Park West , NY 10024-5192, USA. (Ref. 41414) Poll, M.0 Recherches sur la faune ichthyologique de la région du Stanley-Pool. Ann. Mus. R. Congo Belge, Sér. In-8°, Sci. Zool., 71:75-174. (Ref. 41585) David, L. and M. Poll0 Contribution à la faune ichthyologique du Congo Belge: collections du Dr. H. Schouteden (1924-1926) et d'autres récolteurs. Ann. Mus. Congo Belge, Zool. (1), 3(5):189-294. (Ref. 41590) Pellegrin, J.0 Poissons du Chiloango et du Congo recueillis par l'expédition du Dr. Schouteden (1920-1922). Ann. Mus. Congo Belge, Zool. (1), 3(1):1-50. (Ref. 41591) Daget, J. and A. Stauch0 Poissons de la rive droite du Moyen-Congo. Mission A. Stauch (février-avril 1961). Bull. Inst. Rech. Sci. Congo 2:41-48. (Ref. 44840) Gosse, J.-P.0 Les poissons du bassin de l'Ubangi. Mus. R. Afr. Centr., Doc. Zool., 13:56 p. (Ref. 45441) Varjo, M., L. Koli and H. Dahlström0 Kalannimiluettelo (versio 10/03). Suomen Biologian Seura Vanamo Ry. (Ref. 51086) Carl, H.0 Danish fish names. Zoological Museum of Copenhagen. Unpublished. (Ref. 51471) Chinese Academy of Fishery Sciences0 Chinese aquatic germplasm resources database. http://zzzy.fishinfo.cn (Ref. 58108) Norris, S.M.0 Anabantidae. p. 251-268. In M.L.J. Stiassny, G.G. Teugels and C.D. Hopkins (eds.) The fresh and brackish water fishes of Lower Guinea, West-Central Africa. Volume II. Collection Faune et Flore tropicales 42. Institut de Recherche pour le Développement, Paris, France, Muséum National d’Histoire Naturelle, Paris, France, and Musée Royal de l’Afrique Centrale, Tervuren, Belgium. 603 pp. (Ref. 81650) Monsembula Iyaba, R.J.C. and M.L.J. Stiassny0 Fishes of the Salonga National Park (Congo basin, central Africa): a list of species collected in the Luilaka, Salonga, and Yenge Rivers (Equateur Province, Democratic Republic of Congo). Check List 9(2):246-256. (Ref. 93897) Lingofo Bolaya, R.0 Common names of some fish species found in the Isangi Territory (Tshopo District, Orientale Province, Democratic Republic of the Congo). Unpublished compilation of R. Lingofo Bolaya. (Ref. 98754) Moelants, T.0 Diversity and ecology of the ichthyofauna of the Middle and Upper Congo basin: a case-study in the region of the Wagenia falls (Democratic Republic of the Congo). KULeuven, Faculty of Science, Leuven (Belgium), 320p. (Ref. 106245) Decru, E.0 The ichthyofauna in the Central Congo basin: diversity and distribution in the north-eastern tributaries. KULeuven, Faculty of Science, Leuven (Belgium), 303p. (Ref. 106290) Skelton, P.H.0 Chapter 11. The freshwater fishes of Angola. p. 207-242. In B.J. Huntley, V. Russo, F. Lages and N. Ferrand (eds.) Biodiversity of Angola. Science & conservation: a modern synthesis. Springer Nature Switzerland AG, Cham, Switzerland. 549 p. (Ref. 120641) Dean, E.M., A.R. Cooper, L. Wang, W. Daniel, S. David, C. Ernzen, K.B. Gido, E Hale, T.J. Haxton, W. Kelso, N. Leonard, C. Lido, J. Margraf, M. Porter, C. Pennock, D. Propst, J. Ross, M.D. Staudinger, G. Whelan and D.M. Infante0 The North American Freshwater Migratory Fish Database (NAFMFD): Characterizing the migratory life histories of freshwater fishes of Canada, the United States of Mexico. J. Biogeography 49:1193-1203. (Ref. 126106) Stiassny, M.L.J., E. Alter, R.J.C. Monsembula Iyaba and T.L.D. Liyandja0 Two new <i>Phenacogrammus</i> (Characoidei; Alestidae) from the Ndzaa River (Mfimi-Lukenie basin) of central Africa, Democratic Republic of Congo. Am. Mus. Novit. 3980:1-22. (Ref. 127185) Price, S.A., S.T. Friedman, K.A. Corn, O. Larouche, K. Brockelsby, A.J. Lee, M. Nagaraj, N.G. Bertrand, M. Danao, M.C. Coyne, J.R. Estrada, R. Friedman, E. Hoeft, M. Iwan, D. Gross, J.H. Kao, B. Landry, M.J. Linares, C. McGlinn and …0 FishShapes v1: Functionally relevant measurements of teleost shape and size on three dimensions. Ecology 103(e3829):1-2. (Ref. 137577) FB-GeneralDescr-4779 http://purl.org/dc/dcmitype/Text text/html Susan M. Luna 1991-11-28 00:00:00 2025-12-19 00:00:00 Biology http://creativecommons.org/licenses/by-nc/3.0/ WorldFish Center - FishBase General public http://www.fishbase.org/summary/SpeciesSummary.php?id=4779 http://rs.tdwg.org/ontology/voc/SPMInfoItems#Threats Oviparous (Ref. 205). Gosse, J.-P. 1986 Anabantidae. p. 402-414. In J. Daget, J.-P. Gosse and D.F.E. Thys van den Audenaerde (eds.) Check-list of the freshwater fishes of Africa (CLOFFA). ISNB, Brussels; MRAC, Tervuren; and ORSTOM, Paris. Vol. 2. (Ref. 6803) FB-DiagnosticDescr-4779 http://purl.org/dc/dcmitype/Text text/html Tobias Musschoot 2017-10-16 00:00:00 2017-10-16 00:00:00 http://creativecommons.org/licenses/by-nc/3.0/ WorldFish Center - FishBase General public http://www.fishbase.org/summary/SpeciesSummary.php?id=4779 http://rs.tdwg.org/ontology/voc/SPMInfoItems#Habitat Diagnosis: dorsal fin with 18-20 spines (17, 21 rare) and 10-12 (9, 13 rare) soft rays; anal fin with 8-9 spines (10 rare) and 10-12 (9, 13 rare) soft rays; 12-15 (mode 14) pectoral fin rays; dorsal profile rounded; snout blunt; body fusiform; pelvic fin clear; frequently an oblong brown bar on caudal peduncle; overall pigmentation usually grey (Ref. 81650).Description: body typically fusiform, with blunt snout and head; 15-20 scales in upper lateral line, 6-16 in lower; 2-4 scales above upper lateral line, 7-9 below; 2-18 serrations above opercular notch, 0-11 serrations within notch, 2-20 serrations below, 6-20 subopercular serrations, 0-9 interopercular serrations, preopercle entire (Ref. 81650).Coloration: adults dark brownish grey or grey overall, with up to 9 faint bars on flanks; on caudal peduncle 3-4 of these bars fuse into an oblong stripe, a typical feature of juveniles and young adults, but fading with size; membrane in opercular notch frequently black; pelvic fin always clear and unpaired fins not darkly pigmented; juvenile specimens (up to 40 mm SL) have up to 9 black bars on flanks, each one scale row wide (Ref. 81650). Norris, S.M. 2008 Anabantidae. p. 251-268. In M.L.J. Stiassny, G.G. Teugels and C.D. Hopkins (eds.) The fresh and brackish water fishes of Lower Guinea, West-Central Africa. Volume II. Collection Faune et Flore tropicales 42. Institut de Recherche pour le Développement, Paris, France, Muséum National d’Histoire Naturelle, Paris, France, and Musée Royal de l’Afrique Centrale, Tervuren, Belgium. 603 pp. (Ref. 81650) FB-Morphology-4779 http://purl.org/dc/dcmitype/Text text/html Tobias Musschoot 2017-10-16 00:00:00 2017-10-16 00:00:00 http://creativecommons.org/licenses/by-nc/3.0/ WorldFish Center - FishBase General public http://www.fishbase.org/summary/SpeciesSummary.php?id=4779 http://rs.tdwg.org/ontology/voc/SPMInfoItems#Distribution Dorsal spines (total): 17 - 21; Dorsal soft rays (total): 9 - 13; Anal spines: 8 - 10; Analsoft rays: 9 - 13; Vertebrae: 30 - 31 Norris, S.M. 2008 Anabantidae. p. 251-268. In M.L.J. Stiassny, G.G. Teugels and C.D. Hopkins (eds.) The fresh and brackish water fishes of Lower Guinea, West-Central Africa. Volume II. Collection Faune et Flore tropicales 42. Institut de Recherche pour le Développement, Paris, France, Muséum National d’Histoire Naturelle, Paris, France, and Musée Royal de l’Afrique Centrale, Tervuren, Belgium. 603 pp. (Ref. 81650) FB-Distribution-4779 http://purl.org/dc/dcmitype/Text text/html Susan M. Luna 1991-11-28 00:00:00 2022-09-02 00:00:00 http://creativecommons.org/licenses/by-nc/3.0/ WorldFish Center - FishBase General public http://www.fishbase.org/summary/SpeciesSummary.php?id=4779 http://rs.tdwg.org/ontology/voc/SPMInfoItems#Uses Africa: Chiloango and Kouilou basins in Angola (Cabinda Province) and Republic of Congo; also in the Ngongo-Noumbi rivers area in Republic of Congo (Ref. 81650). Common throughout most of the Congo River basin (Ref. 1880, 41585, 41591, 45441, 81650, 93897, 106245, 106290). Report from the Ogowe River (Ref. 1880) unconfirmed (Ref. 81650). Widely transported by the aquarium fish trade. Gosse, J.-P. 1986 Anabantidae. p. 402-414. In J. Daget, J.-P. Gosse and D.F.E. Thys van den Audenaerde (eds.) Check-list of the freshwater fishes of Africa (CLOFFA). ISNB, Brussels; MRAC, Tervuren; and ORSTOM, Paris. Vol. 2. (Ref. 6803) FB-Uses-4779 http://purl.org/dc/dcmitype/Text text/html Susan M. Luna 1991-11-28 00:00:00 2025-12-19 00:00:00 Importance http://creativecommons.org/licenses/by-nc/3.0/ WorldFish Center - FishBase General public http://www.fishbase.org/summary/SpeciesSummary.php?id=4779 http://rs.tdwg.org/ontology/voc/SPMInfoItems#LifeCycle aquarium: commercial Baensch, H.A. and R. Riehl 1985 Aquarien atlas. Band 2. Mergus, Verlag für Natur-und Heimtierkunde GmbH, Melle, Germany. 1216 p. (Ref. 2060) FB-LifeCycle-4779 http://purl.org/dc/dcmitype/Text text/html Susan M. Luna 1991-11-28 00:00:00 2025-12-19 00:00:00 http://creativecommons.org/licenses/by-nc/3.0/ WorldFish Center - FishBase General public http://www.fishbase.org//Reproduction/FishReproSummary.php?ID=4779&StockCode=5003 http://rs.tdwg.org/ontology/voc/SPMInfoItems#TrophicStrategy Oviparous (Ref. 205). Breder, C.M. and D.E. Rosen 1966 Modes of reproduction in fishes. T.F.H. Publications, Neptune City, New Jersey. 941 p. (Ref. 205) FB-Size-4779 http://purl.org/dc/dcmitype/Text text/html Susan M. Luna 1991-11-28 00:00:00 2025-12-19 00:00:00 Max. size http://creativecommons.org/licenses/by-nc/3.0/ WorldFish Center - FishBase General public http://www.fishbase.org/summary/SpeciesSummary.php?id=4779 http://rs.tdwg.org/ontology/voc/SPMInfoItems#Size 17.0 cm TL (male/unsexed; (Ref. 6803)) Gosse, J.-P. 1986 Anabantidae. p. 402-414. In J. Daget, J.-P. Gosse and D.F.E. Thys van den Audenaerde (eds.) Check-list of the freshwater fishes of Africa (CLOFFA). ISNB, Brussels; MRAC, Tervuren; and ORSTOM, Paris. Vol. 2. (Ref. 6803) FB-Threats-2 http://purl.org/dc/dcmitype/Text text/html Belen Acosta 1990-10-18 00:00:00 2024-03-07 00:00:00 http://creativecommons.org/licenses/by-nc/3.0/ WorldFish Center - FishBase General public http://www.fishbase.org/summary/SpeciesSummary.php?id=2 http://rs.tdwg.org/ontology/voc/SPMInfoItems#Threats Least Concern (LC) , IUCN Grouper and Wrasse Specialist Group IUCN 2006 2006 IUCN red list of threatened species. www.iucnredlist.org. Downloaded July 2006. FB-Habitat-4779 http://purl.org/dc/dcmitype/Text text/html Susan M. Luna 1991-11-28 00:00:00 2025-12-19 00:00:00 Environment http://creativecommons.org/licenses/by-nc/3.0/ WorldFish Center - FishBase General public http://www.fishbase.org/summary/SpeciesSummary.php?id=4779 http://rs.tdwg.org/ontology/voc/SPMInfoItems#Diseases benthopelagic; non-migratory (Ref. 126106); freshwater Dean, E.M., A.R. Cooper, L. Wang, W. Daniel, S. David, C. Ernzen, K.B. Gido, E Hale, T.J. Haxton, W. Kelso, N. Leonard, C. Lido, J. Margraf, M. Porter, C. Pennock, D. Propst, J. Ross, M.D. Staudinger, G. Whelan and D.M. Infante 2022 The North American Freshwater Migratory Fish Database (NAFMFD): Characterizing the migratory life histories of freshwater fishes of Canada, the United States of Mexico. J. Biogeography 49:1193-1203. (Ref. 126106)