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<taxon>
	<dc:identifier>FB-4483</dc:identifier>
	<dc:source>http://www.fishbase.org/summary/SpeciesSummary.php?id=4483</dc:source>
	<dwc:Kingdom>Animalia</dwc:Kingdom>
	<dwc:Phylum>Chordata</dwc:Phylum>
	<dwc:Class>Teleostei</dwc:Class>
	<dwc:Order>Cypriniformes</dwc:Order>
	<dwc:Family>Acheilognathidae</dwc:Family>
	<dwc:Genus>Rhodeus</dwc:Genus>
	<dwc:ScientificName>Rhodeus amarus (Bloch, 1782)</dwc:ScientificName>
	<commonName xml:lang='fa'>Mahi Mokhrj Lulehai</commonName>
	<commonName xml:lang='fa'>Mahi-e-Makhraj looleei</commonName>
	<commonName xml:lang='pl'>Rózanka europejska</commonName>
	<commonName xml:lang='sq'>Idhtaku</commonName>
	<commonName xml:lang='cs'>Hořavka hořká</commonName>
	<commonName xml:lang='da'>Bitterling</commonName>
	<commonName xml:lang='da'>Europæisk bitterling</commonName>
	<commonName xml:lang='de'>Bauerkärpflein</commonName>
	<commonName xml:lang='de'>Bille</commonName>
	<commonName xml:lang='de'>Bitterfisch</commonName>
	<commonName xml:lang='de'>Bitterling</commonName>
	<commonName xml:lang='de'>Blaukärpfli</commonName>
	<commonName xml:lang='de'>Blecke</commonName>
	<commonName xml:lang='de'>Burghoferl</commonName>
	<commonName xml:lang='de'>Kobauch</commonName>
	<commonName xml:lang='de'>Lieschkarpfen</commonName>
	<commonName xml:lang='de'>Paffendume</commonName>
	<commonName xml:lang='de'>Pille</commonName>
	<commonName xml:lang='de'>Plätteln</commonName>
	<commonName xml:lang='de'>Plättken</commonName>
	<commonName xml:lang='de'>Schbneiderkarpfen</commonName>
	<commonName xml:lang='de'>Schniederkarpfen</commonName>
	<commonName xml:lang='de'>Weberle</commonName>
	<commonName xml:lang='de'>Wittfisch</commonName>
	<commonName xml:lang='en'>Bitterling</commonName>
	<commonName xml:lang='en'>European bitterling</commonName>
	<commonName xml:lang='et'>Euroopa mõrukas</commonName>
	<commonName xml:lang='fi'>Katkerokala</commonName>
	<commonName xml:lang='fr'>Bitterling</commonName>
	<commonName xml:lang='fr'>Bouvière</commonName>
	<commonName xml:lang='fr'>Péteuse</commonName>
	<commonName xml:lang='fr'>Rosière</commonName>
	<commonName xml:lang='hu'>szivárványos ökle</commonName>
	<commonName xml:lang='ru'>Gorchak</commonName>
	<commonName xml:lang='ru'>европейский обыкновенный горчак</commonName>
	<commonName xml:lang='tr'>Acıbalık</commonName>
	<commonName xml:lang=''>Behlita</commonName>
	<commonName xml:lang=''>Blehnita</commonName>
	<commonName xml:lang=''>Boarca</commonName>
	<commonName xml:lang=''>Boarta</commonName>
	<commonName xml:lang=''>Bortanica</commonName>
	<commonName xml:lang=''>Burtica</commonName>
	<commonName xml:lang=''>Chisoaga</commonName>
	<commonName xml:lang=''>Mourmouritsa</commonName>
	<commonName xml:lang=''>Pezdirk</commonName>
	<commonName xml:lang=''>Plutica</commonName>
	<commonName xml:lang=''>Βαβούκι</commonName>
	<commonName xml:lang=''>Κοκκινομάτης</commonName>
	<commonName xml:lang=''>Μουρμουρίτσα</commonName>
	<commonName xml:lang=''>Φλασκούνι</commonName>
	<commonName xml:lang=''>苦味鰟鲏</commonName>
	<commonName xml:lang=''>苦味鳑鲏</commonName>
	<synonym relationship='synonym'>Cyprinus amarus Bloch, 1782</synonym>
	<synonym relationship='misapplied name'>Rhodeus amarus meridionalis (non Karaman, 1924)</synonym>
	<synonym relationship='synonym'>Rhodeus genitalis Walecki, 1863</synonym>
	<synonym relationship='synonym'>Rhodeus lucinae Walecki, 1863</synonym>
	<synonym relationship='misapplied name'>Rhodeus sericeus (non Pallas, 1776)</synonym>
	<synonym relationship='synonym'>Rhodeus sericeus amarus (Bloch, 1782)</synonym>
	<dcterms:created>1991-06-21</dcterms:created>
	<dcterms:modified>2016-05-02</dcterms:modified>
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	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=58297&amp;speccode=4483'>Bogutskaya, N.G. and A.M. Naseka0 REGIONAL CHECK-LISTS: Volga River Drainage Area. In Website and Database: "Freshwater Fishes of Russia": A Source of Information on the Current State of the Fauna. Zoological Institute RAS. (Ref. 58297)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=58342&amp;speccode=4483'>Fricke, R., M. Bilecenoglu and H.M. Sari0 Annotated checklist of fish and lamprey species (Gnathostoma and Petromyzontomorphi) of Turkey, including a Red List of threatened and declining species. Stuttgarter Beitr. Naturk. Sea A (706):1-172. (Ref. 58342)</reference>
	<reference isbn='978-2-8399-0298-4' url='http://www.fishbase.org/references/FBRefSummary.php?id=59043&amp;speccode=4483'>Kottelat, M. and J. Freyhof0 Handbook of European freshwater fishes. Publications Kottelat, Cornol and Freyhof, Berlin. 646 pp. (Ref. 59043)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=59105&amp;speccode=4483'>Tarkan, A.S., O. Gaygusuz, H. Acipinar, C. Gürsoy and M. Ozulug0 Length-weight relationship of fishes from the Marmara region (NW-Turkey). J. Appl. Ichthyol. 22:271-273. (Ref. 59105)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=74657&amp;speccode=4483'>Bartley, D.M. (comp./ed.)0 DIAS. Introduced species in fisheries and aquaculture: information for responsible use and control. CD-ROM. Rome (Italy): FAO. (Ref. 74657)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=75668&amp;speccode=4483'>Crivelli, A.J., G. Catsadorakis, M. Malakou and E. Rosecchi0 Fish and fisheries of the Prespa lakes. Hydrobiologia 351:107-125. (Ref. 75668)</reference>
	<reference isbn='978-99956-02-70-3' url='http://www.fishbase.org/references/FBRefSummary.php?id=83428&amp;speccode=4483'>Dhora, Dh.0 Regjistër I specieve të faunës së Shqipërisë 2010. Shkodër: Camaj-Pipa. 208 p. (Ref. 83428)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=83882&amp;speccode=4483'>FAO-FIES0 Aquatic Sciences and Fisheries Information System (ASFIS) species list. Retrieved from http://www.fao.org/fishery/collection/asfis/en, March 2010. (Ref. 83882)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=86351&amp;speccode=4483'>Özcan, G.0 The status of the freshwater gobiid, &lt;i&gt;Knipowitschia mermere&lt;/i&gt; Ahnelt, 1995: distribution ecology and threats. Biharean Biol. 3(2):139-141. (Ref. 86351)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=87029&amp;speccode=4483'>Dhora, Dh.0 Fjalor i emrave të kafshëve të Shqipërisë (Emri shkencor - Shqip - Anglisht) / Dictionary of animal names of Albania (Scientific names - Albanian - English). Camaj - Pipa. Shkodër, Albania. pp. 288. (Ref. 87029)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=87703&amp;speccode=4483'>AFMAFEWM (Austrian Federal Ministry of Agriculture, Forestry, Environment and Water Management)0 Checklist of species in Austria. Federal Office for Water Management, Institute of Waters Ecology, Fisheries and Lake Research. (Ref. 87703)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=88166&amp;speccode=4483'>Verreycken, H., G. Van Thuyne and C. Belpaire0 Length-weight relationships of 40 freshwater fish species from two decades of monitoring in Flanders (Belgium). J. Appl. Ichthyol. 2011:1-5. (Ref. 88166)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=89840&amp;speccode=4483'>Lusk, S., V. Luskova, L. Hanel, B. Lojkasek and P. Hartvich0 Červený seznam mihulí a ryb České Republiky - verze 2010. The Red List of lampreys and fishes of the Czech Republic Version 2010. Biodiverzita Ichtyofauny ČR (VIII): 68-78. (Ref. 89840)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=90061&amp;speccode=4483'>Povz, M.0 The red data list of the freshwater lampreys (Cyclostomata) and fish (Pisces) of Slovenia. p. 63-72. A. Kirchhofer and D. Hefti (eds) In Conservation of Endangered Freshwater Fish in Europe. Birkhäuser Verlag, Basel, Switzerland. (Ref. 90061)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=90062&amp;speccode=4483'>FAO-FIES0 Aquatic Sciences and Fisheries Information System (ASFIS) species list. Retrieved from http://www.fao.org/fishery/collection/asfis/en, March 2012. (Ref. 90062)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=94493&amp;speccode=4483'>Bianco, P.G.0 An update on the status of native and exotic freshwater fishes of Italy. J. Appl. Ichthyol. 30(1):62-77. (Ref. 94493)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=94844&amp;speccode=4483'>Ilhan, A., M. Sari, . Saygi and M.R. Ustaoğlu0 Length-weight relationships of freshwater fishes in the Biga Peninsula (northwestern Anatolia, Turkey). J. Appl. Ichthyol. 28:857-858. (Ref. 94844)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=95340&amp;speccode=4483'>Petriki, O., E. Gousia and D.C. Bobori0 Weight-length relationships of 36 fish species from the River Strymon system (northern Greece). J. Appl. Ichthyol. 27(3):939-941. (Ref. 95340)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=95632&amp;speccode=4483'>FAO-FIES0 Aquatic Sciences and Fisheries Information System (ASFIS) species list. Retrieved from http://www.fao.org/fishery/collection/asfis/en, April 2014. (Ref. 95632)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=101110&amp;speccode=4483'>FAO-FIES0 Aquatic Sciences and Fisheries Information System (ASFIS) species list. Retrieved from http://www.fao.org/fishery/collection/asfis/en, [accessed 13/04/2015]. (Ref. 101110)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=105471&amp;speccode=4483'>eElurikkus0 eElurikkus koondab ühte portaali Eesti eluslooduse andmed. http://elurikkus.ut.ee/ (Ref. 105471)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=106151&amp;speccode=4483'>Çiçek, E., R. Fricke and S.S. Birecikligil0 Freshwater fishes of Turkey: a revised and updated annotated checklist. Biharean Biologist 9(2):141-157. (Ref. 106151)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=115257&amp;speccode=4483'>FAO-FIES0 Aquatic Sciences and Fisheries Information System (ASFIS) species list. Retrieved from http://www.fao.org/fishery/collection/asfis/en (accessed 08/06/2017). (Ref. 115257)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=118272&amp;speccode=4483'>FAO-FIES0 Aquatic Sciences and Fisheries Information System (ASFIS) species list. Retrieved from htttps://www.fao.org/fishery/collection/asfis/en [accessed May, 2018] (Ref. 118272)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=121524&amp;speccode=4483'>Mouladi-Saleh, A. and S. Eagderi0 Length-weight relationship and condition factor of ten fish species (Cyprinidae, Sisoridae, Mugilidae, Cichlidae, Gobiidae and Channidae) from Iranian inland waters. J. Wildlife Bio. 3(4):12-15. (Ref. 121524)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=125501&amp;speccode=4483'>Dhora, D.0 Updated lists of the fish species of Albania's freshwater. Bul. Shk. Ser. Shk. Nat. 2020(70):46-74. (Ref. 125501)</reference>
	<reference isbn='2350-4757' url='http://www.fishbase.org/references/FBRefSummary.php?id=128520&amp;speccode=4483'>Marčeta, B. and K. Pliberšek (eds)0 BiosWeb. Biological database of the Fisheries Research Institute of Slovenia. Ljubljana, Fisheries Research Institute of Slovenia, 2014. (www.biosweb.org. updated 13/04/2023). (Ref. 128520)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=130435&amp;speccode=4483'>IUCN0 The IUCN Red List of Threatened Species. 1964 - onwards. https://www.iucnredlist.org/ (Ref. 130435)</reference><dataObject>
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	<dc:description>Occurs most abundantly in still or slow-flowing water with dense aquatic vegetation and sand-silt bottom as lowland ponds, canals, slow-flowing rivers, backwaters and oxbows, where mussels are present (Ref. 59043).  Found among plants over sand and muddy bottoms in shallow waters.  Remarkable for its habit of deposing its eggs in the cavity of bivalves (Unio).  Feeds mainly on plants and to a lesser degree on worms, crustaceans, and insect larvae.  Formerly used for pregnancy tests:  females injected with urine from pregnant women protruded their ovipositors (Ref. 6258).  Live span is exceptionally up to 5 years but most individuals do not survive the year of their first reproduction and population sizes fluctuate  greatly over the years (Ref. 59043).</dc:description>
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	<reference isbn='978-2-8399-0298-4' url='http://www.fishbase.org/references/FBRefSummary.php?id=59043&amp;speccode=4483'>Kottelat, M. and J. Freyhof 2007 Handbook of European freshwater fishes. Publications Kottelat, Cornol and Freyhof, Berlin. 646 pp. (Ref. 59043)</reference></dataObject><dataObject>
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	<dc:description>Caudal fin with 19 to 20 soft rays.  Differs from Rhodeus meridionalis by having sub-inferior mouth (vs. terminal), rostral cap covering all or at least more than half of upper lip (vs. only upper part of upper lip) (Ref. 59043).</dc:description>
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	<reference isbn='2-85653653261' url='http://www.fishbase.org/references/FBRefSummary.php?id=40476&amp;speccode=4483'>Keith, P. and J. Allardi (coords.) 2001 Atlas des poissons d'eau douce de France. Muséum national d’Histoire naturelle, Paris. Patrimoines naturels, 47:1-387. (Ref. 40476)</reference></dataObject><dataObject>
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	<dc:description>Dorsal spines (total): 3; Dorsal soft rays (total): 8 - 10;  Analspines: 3; Analsoft rays: 8 - 10; Vertebrae: 34 - 36</dc:description>
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	<reference isbn='2-85653653261' url='http://www.fishbase.org/references/FBRefSummary.php?id=40476&amp;speccode=4483'>Keith, P. and J. Allardi (coords.) 2001 Atlas des poissons d'eau douce de France. Muséum national d’Histoire naturelle, Paris. Patrimoines naturels, 47:1-387. (Ref. 40476)</reference></dataObject><dataObject>
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	<dc:description>Europe:  central and eastern Europe and northern Asia Minor (Ref. 57778).  Basins of North, southern Baltic, Black, western and southern Caspian and Aegean Seas (from Maritza to Struma drainages); Mediterranean basin, only in northern Rhône (France) and Drin drainages (Alabania, Montenegro,  Macedonia).  Abundant and expanding in most of its range, but locally threatened by water pollution, weed clearing, and stocking of predatory fish (Ref. 59043).</dc:description>
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	<reference isbn='978-2-8399-0298-4' url='http://www.fishbase.org/references/FBRefSummary.php?id=59043&amp;speccode=4483'>Kottelat, M. and J. Freyhof 2007 Handbook of European freshwater fishes. Publications Kottelat, Cornol and Freyhof, Berlin. 646 pp. (Ref. 59043)</reference></dataObject><dataObject>
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	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=7322&amp;speccode=4483'>Ladiges, W. and D. Vogt 1979 Die Süsswasserfische Europas. Paul Parey, Hamburg. 299 p. (Ref. 7322)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=7322&amp;speccode=4483'>Ladiges, W. and D. Vogt 1979 Die Süsswasserfische Europas. Paul Parey, Hamburg. 299 p. (Ref. 7322)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=6258&amp;speccode=4483'>Maitland, P.S. and R.N. Campbell 1992 Freshwater fishes of the British Isles. HarperCollins Publishers, London.368 p. (Ref. 6258)</reference></dataObject><dataObject>
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	<dc:description>Spawns in clear, slow-running or still water, often with a muddy bottom. The presence of nearby freshwater mussels is of vital importance. Female deposits eggs within the valves of the mussel. Male sheds sperm into the inhalent current of the respiring mussel and thereby, fertilizes the eggs. Fry remain in the mussel until protection is not anymore required.</dc:description>
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	<reference isbn='0-600-33476-7.' url='http://www.fishbase.org/references/FBRefSummary.php?id=9696&amp;speccode=4483'>Vostradovsky, J. 1973 Freshwater fishes. The Hamlyn Publishing Group Limited, London. 252 p. (Ref. 9696)</reference></dataObject><dataObject>
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	<agent homepage='' logoURL='' role='recorder'>Drina Sta. Iglesia</agent>
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	<dc:source>amarus</dc:source>
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	<dc:description>Among plants over sand and muddy bottoms in shallow waters. Remarkable for its habit of deposing its eggs in the cavity of bivalves (Unio). Feeds mainly on plants and to a lesser degree on worms, crustaceans, and insect larvae.</dc:description>
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	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=13696&amp;speccode=4483'>Kottelat, M. 1997 European freshwater fishes. An heuristic checklist of the freshwater fishes of Europe (exclusive of former USSR), with an introduction for non-systematists and comments on nomenclature and conservation. Biologia, Bratislava, 52/Suppl. 5:1-271. (Ref. 13696)</reference></dataObject><dataObject>
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	<dc:description>11.2 cm TL (male/unsexed; (Ref. 88166)); max. reported age: 5 years (Ref. 9696)</dc:description>
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	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=88166&amp;speccode=4483'>Verreycken, H., G. Van Thuyne and C. Belpaire 2011 Length-weight relationships of 40 freshwater fish species from two decades of monitoring in Flanders (Belgium). J. Appl. Ichthyol. 2011:1-5. (Ref. 88166)</reference>
	<reference isbn='0-600-33476-7.' url='http://www.fishbase.org/references/FBRefSummary.php?id=9696&amp;speccode=4483'>Vostradovsky, J. 1973 Freshwater fishes. The Hamlyn Publishing Group Limited, London. 252 p. (Ref. 9696)</reference></dataObject><dataObject>
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	<dc:source>http://www.fishbase.org/summary/SpeciesSummary.php?id=2</dc:source>
	<subject>http://rs.tdwg.org/ontology/voc/SPMInfoItems#Distribution</subject>
	<dc:description>Least Concern (LC) , IUCN Grouper and Wrasse Specialist Group</dc:description>
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	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=57073'>IUCN 2006 2006 IUCN red list of threatened species. www.iucnredlist.org. Downloaded July 2006.</reference></dataObject><dataObject>
	<dc:identifier>FB-Habitat-4483</dc:identifier>
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	<agent homepage='' logoURL='' role='recorder'>Crispina B. Binohlan</agent>
	<dcterms:created>1991-06-21 00:00:00</dcterms:created>
	<dcterms:modified>2016-05-02 00:00:00</dcterms:modified>
	<dc:title>Environment</dc:title>
	<dc:language></dc:language>
	<license>http://creativecommons.org/licenses/by-nc/3.0/</license>
	<dc:rights></dc:rights>
	<dcterms:bibliographicCitation></dcterms:bibliographicCitation>
	<dc:source>http://www.fishbase.org/summary/SpeciesSummary.php?id=4483</dc:source>
	<dc:description>benthopelagic; freshwater; depth range 0 - ? m</dc:description>
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