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<taxon>
	<dc:identifier>FB-4480</dc:identifier>
	<dc:source>http://www.fishbase.org/summary/SpeciesSummary.php?id=4480</dc:source>
	<dwc:Kingdom>Animalia</dwc:Kingdom>
	<dwc:Phylum>Chordata</dwc:Phylum>
	<dwc:Class>Petromyzonti</dwc:Class>
	<dwc:Order>Petromyzontiformes</dwc:Order>
	<dwc:Family>Petromyzontidae</dwc:Family>
	<dwc:Genus>Lampetra</dwc:Genus>
	<dwc:ScientificName>Lampetra fluviatilis (Linnaeus, 1758)</dwc:ScientificName>
	<commonName xml:lang='no'>Elveniøye</commonName>
	<commonName xml:lang='pl'>Minog</commonName>
	<commonName xml:lang='pl'>Minog rzeczny</commonName>
	<commonName xml:lang='pl'>Minóg rzeczny</commonName>
	<commonName xml:lang='pt'>Lampreia do rio</commonName>
	<commonName xml:lang='pt'>Lampreia-de-rio</commonName>
	<commonName xml:lang='pt'>Lampreia-do-rio</commonName>
	<commonName xml:lang='pt'>Lampreira-de-rio</commonName>
	<commonName xml:lang='sq'>Kavall lumi</commonName>
	<commonName xml:lang='cs'>Mihula riecna</commonName>
	<commonName xml:lang='cs'>Mihule potocní</commonName>
	<commonName xml:lang='cs'>Mihule říční</commonName>
	<commonName xml:lang='cy'>Llysywen bendoll yr afon</commonName>
	<commonName xml:lang='da'>Almindelig flodlampret</commonName>
	<commonName xml:lang='da'>Flodlampret</commonName>
	<commonName xml:lang='da'>Flodnegenøjn</commonName>
	<commonName xml:lang='da'>Flodniøje</commonName>
	<commonName xml:lang='de'>Bär</commonName>
	<commonName xml:lang='de'>Bärel</commonName>
	<commonName xml:lang='de'>Flußneunauge</commonName>
	<commonName xml:lang='de'>Flußpricke</commonName>
	<commonName xml:lang='de'>Gemeines</commonName>
	<commonName xml:lang='de'>Lamprete</commonName>
	<commonName xml:lang='de'>Lamprida</commonName>
	<commonName xml:lang='de'>Lampride</commonName>
	<commonName xml:lang='de'>Nägenooge</commonName>
	<commonName xml:lang='de'>Negenauge</commonName>
	<commonName xml:lang='de'>Negenooge</commonName>
	<commonName xml:lang='de'>NeünAug</commonName>
	<commonName xml:lang='de'>Neunauge</commonName>
	<commonName xml:lang='de'>Pricke</commonName>
	<commonName xml:lang='de'>Querder</commonName>
	<commonName xml:lang='de'>Rheinpricke</commonName>
	<commonName xml:lang='de'>Uhlen</commonName>
	<commonName xml:lang='en'>European river lamprey</commonName>
	<commonName xml:lang='en'>Lampern</commonName>
	<commonName xml:lang='en'>River lamprey</commonName>
	<commonName xml:lang='en'>Stone eel</commonName>
	<commonName xml:lang='et'>Jõesilm</commonName>
	<commonName xml:lang='et'>Silmus</commonName>
	<commonName xml:lang='fi'>flodnejonöga</commonName>
	<commonName xml:lang='fi'>Nahkiainen</commonName>
	<commonName xml:lang='fr'>Châtillon</commonName>
	<commonName xml:lang='fr'>Filtre</commonName>
	<commonName xml:lang='fr'>Flûte</commonName>
	<commonName xml:lang='fr'>Lamprillon</commonName>
	<commonName xml:lang='fr'>Lamproie de rivière</commonName>
	<commonName xml:lang='fr'>Lamproie de rivière européene</commonName>
	<commonName xml:lang='fr'>Lamproie fluviatile</commonName>
	<commonName xml:lang='fr'>Lamproie fluviatile européene</commonName>
	<commonName xml:lang='fr'>Sept-trous</commonName>
	<commonName xml:lang='is'>Fisksuga</commonName>
	<commonName xml:lang='it'>Lampreda di fiume</commonName>
	<commonName xml:lang='lv'>strauta nēģis</commonName>
	<commonName xml:lang='lt'>upinė nėgė</commonName>
	<commonName xml:lang='mk'>Zmiulka</commonName>
	<commonName xml:lang='ru'>Baltiyskaya minoga</commonName>
	<commonName xml:lang='ru'>Evropeiskaya rechnaya minoga</commonName>
	<commonName xml:lang='ru'>Retschnaja</commonName>
	<commonName xml:lang='ru'>Retschnaja minoga</commonName>
	<commonName xml:lang='ru'>Минога речная</commonName>
	<commonName xml:lang='ru'>минога речная</commonName>
	<commonName xml:lang='ru'>речная минога</commonName>
	<commonName xml:lang='sr'>Paklara rijecna</commonName>
	<commonName xml:lang='sr'>Paklara riječna</commonName>
	<commonName xml:lang='sr'>Rijecna paklara</commonName>
	<commonName xml:lang='sk'>Mihula potocná</commonName>
	<commonName xml:lang='sv'>Flodnejonöga</commonName>
	<commonName xml:lang='tr'>Bofa balığı</commonName>
	<commonName xml:lang='tr'>Bofa balýg¢ý</commonName>
	<commonName xml:lang='tr'>Dere bofa balığı</commonName>
	<commonName xml:lang='es'>Lamprea de río</commonName>
	<commonName xml:lang=''>Amproye</commonName>
	<commonName xml:lang=''>Chiscar de râu</commonName>
	<commonName xml:lang=''>Llampresa de riu</commonName>
	<commonName xml:lang=''>Peist da shuil deag</commonName>
	<commonName xml:lang=''>Potamolamprena</commonName>
	<commonName xml:lang=''>Recni piskur</commonName>
	<commonName xml:lang=''>Rečni piškur</commonName>
	<commonName xml:lang=''>Rivierprik</commonName>
	<commonName xml:lang=''>Ποταμολάμπραινα</commonName>
	<commonName xml:lang=''>七鰓鰻</commonName>
	<commonName xml:lang=''>七鳃鳗</commonName>
	<synonym relationship='synonym'>Ammocoetes communis Gistel, 1848</synonym>
	<synonym relationship='misapplied name'>Lampetra ayresi (non Günther, 1870)</synonym>
	<synonym relationship='ambiguous synonym'>Lampetra fluviatilis f. praecox Berg, 1932</synonym>
	<synonym relationship='ambiguous synonym'>Lampetra fluviatilis ladogensis Ivanova-Berg, 1966</synonym>
	<synonym relationship='synonym'>Lampetra fluviatilis typica Berg, 1931</synonym>
	<synonym relationship='synonym'>Lampetra opisthodon Gratzianov, 1907</synonym>
	<synonym relationship='synonym'>Petromyzon argenteus Bloch, 1795</synonym>
	<synonym relationship='ambiguous synonym'>Petromyzon argenteus Nardo, 1847</synonym>
	<synonym relationship='synonym'>Petromyzon branchialis Linnaeus, 1758</synonym>
	<synonym relationship='synonym'>Petromyzon fluviatilis Linnaeus, 1758</synonym>
	<synonym relationship='ambiguous synonym'>Petromyzon fluviatilis f. major Smitt, 1895</synonym>
	<synonym relationship='synonym'>Petromyzon jurae MacCulloch, 1819</synonym>
	<synonym relationship='ambiguous synonym'>Petromyzon macrops Blainville, 1825</synonym>
	<synonym relationship='synonym'>Petromyzon omalii Beneden, 1857</synonym>
	<synonym relationship='ambiguous synonym'>Petromyzon pricka Lacepède, 1798</synonym>
	<synonym relationship='synonym'>Petromyzon sanguisuga Lacepède, 1800</synonym>
	<dcterms:created>1991-06-21</dcterms:created>
	<dcterms:modified>2024-01-23</dcterms:modified>
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	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=85049&amp;speccode=4480'>Brylińska, M.0 Ryby slodkowodne Polski. PWN, Warszawa. Pp. 521. (Ref. 85049)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=89241&amp;speccode=4480'>Renaud, C.B.0 Lampreys of the world. An annotated and illustrated catalogue of lamprey species known to date. FAO Species Catalogue for Fishery Purposes. No. 5. Rome, FAO. 109 pp. (Ref. 89241)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=89370&amp;speccode=4480'>Lang, N.J., K.J. Roe, C.B. Renaud, H.S. Gill, I.C. Potter, J. Freyhof, A.M. Naseka, P. Cochran, H.E. Pérez, E.M. Habit, B.R. Kuhajda, D.A. Neely, Y.S. Reshetnikov, V.B. Salnikov, M.Th. Stoumboudi and R.L. Mayden0 Novel relationships among lampreys (Petromyzontiformes) revealed by a taxonomically comprehensive molecular data set. Am. Fish. Soc. Symp. 72:41-55. (Ref. 89370)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=89840&amp;speccode=4480'>Lusk, S., V. Luskova, L. Hanel, B. Lojkasek and P. Hartvich0 Červený seznam mihulí a ryb České Republiky - verze 2010. The Red List of lampreys and fishes of the Czech Republic Version 2010. Biodiverzita Ichtyofauny ČR (VIII): 68-78. (Ref. 89840)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=90052&amp;speccode=4480'>Maitland, P.S. and A.A. Lyle0 Threatened freshwater fishes of Great Britain. p. 9-21. A. Kirchhofer and D. Hefti (eds) In Conservation of Endangered Freshwater Fish in Europe. Birkhäuser Verlag, Basel, Switzerland. (Ref. 90052)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=90060&amp;speccode=4480'>Lusk, S.0 The status of the fish fauna in the Czech Republic. p. 89-98. A. Kirchhofer and D. Hefti (eds) In Conservation of Endangered Freshwater Fish in Europe. Birkhäuser Verlag, Basel, Switzerland. (Ref. 90060)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=90062&amp;speccode=4480'>FAO-FIES0 Aquatic Sciences and Fisheries Information System (ASFIS) species list. Retrieved from http://www.fao.org/fishery/collection/asfis/en, March 2012. (Ref. 90062)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=94493&amp;speccode=4480'>Bianco, P.G.0 An update on the status of native and exotic freshwater fishes of Italy. J. Appl. Ichthyol. 30(1):62-77. (Ref. 94493)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=95489&amp;speccode=4480'>Goren, M.0 The fishes of the Mediterranean: a biota under siege. pp. 385-400. In Goffredo, S. and Z. Dubinsky. 2014. The Mediterranean Sea: its history and present challenges. Springer Netherlands. 655 pp. (Ref. 95489)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=95632&amp;speccode=4480'>FAO-FIES0 Aquatic Sciences and Fisheries Information System (ASFIS) species list. Retrieved from http://www.fao.org/fishery/collection/asfis/en, April 2014. (Ref. 95632)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=95727&amp;speccode=4480'>Lloris, D., J. Rucabado, Ll. del Cerro, F. Portas, M. Demestre and A. Roig0 Tots els peixos del Mar Català. I: Llistat de cites I referències. Treballs Soc. Cat. Ict. Herp. 1:1-208. (Ref. 95727)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=95871&amp;speccode=4480'>Psomadakis, P.N., S. Giustino and M. Vacchi0 Mediterranean fish biodiversity: an updated inventory with focus on the Ligurian and Tyrrhenian seas. Zootaxa 3263:1-46. (Ref. 95871)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=101110&amp;speccode=4480'>FAO-FIES0 Aquatic Sciences and Fisheries Information System (ASFIS) species list. Retrieved from http://www.fao.org/fishery/collection/asfis/en, [accessed 13/04/2015]. (Ref. 101110)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=105472&amp;speccode=4480'>Pihu, E.0 Loomade elu 4. köide. Kalad. Pihu, E. 1979. Loomade Elu, 4. Köide, Kalad. Valgus, Tallinn. (Ref. 105472)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=106276&amp;speccode=4480'>Wilhelms, I.0 Atlas of length-weight relationships of 93 fish and crustacean species from the North Sea and the North-East Atlantic (No. 12). Johann Heinrich von Thünen Institute, Federal Research Institute for Rural Areas, Forestry and Fisheries, 552 p. (Ref. 106276)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=115257&amp;speccode=4480'>FAO-FIES0 Aquatic Sciences and Fisheries Information System (ASFIS) species list. Retrieved from http://www.fao.org/fishery/collection/asfis/en (accessed 08/06/2017). (Ref. 115257)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=116450&amp;speccode=4480'>Béarez, P., P. Pruvost, É. Feunteun, S. Iglésias, P. Francour, R. Causse, J. De Mazières, S. Tercerie and N. Bailly0 Checklist of the marine fishes from metropolitan France. Cybium 41(4):351-371. (Ref. 116450)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=118521&amp;speccode=4480'>Salas, C.A., K.E. Yopak, T.J. Lisney, I.C. Potter and S.P. Collin0 The central nervous system of jawless vertebrates: encephalization in lampreys and hagfishes. Brain Behav. Evol. (Ref. 118521)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=119049&amp;speccode=4480'>Antunes, C., F. Cobo and M.J. Araujo0 Iberian inland fisheries. p. 268-282. In J.F. Craig (ed.).  Freshwater Fisheries Ecology. John Wiley &amp; Sons Ltd. 899p. (Ref. 119049)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=119201&amp;speccode=4480'>Bianco, P.G. and V. Ketmaier0 Nature and status of freshwater and estuarine fisheries in Italy and Western Balkans. p. 283-291. In J.F. Craig (ed.).  Freshwater Fisheries Ecology. John Wiley &amp; Sons Ltd. 899p. (Ref. 119201)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=128458&amp;speccode=4480'>Martins, R. and M. Carneiro0 Manual de identificação de peixes ósseos da costa continental portuguesa. Principais características diagnosticantes. IPMA, Instituto Português do Mar e da Atmosfera, Lisboa:1-204. (Ref. 128458)</reference>
	<reference isbn='2350-4757' url='http://www.fishbase.org/references/FBRefSummary.php?id=128520&amp;speccode=4480'>Marčeta, B. and K. Pliberšek (eds)0 BiosWeb. Biological database of the Fisheries Research Institute of Slovenia. Ljubljana, Fisheries Research Institute of Slovenia, 2014. (www.biosweb.org. updated 13/04/2023). (Ref. 128520)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=130435&amp;speccode=4480'>IUCN0 The IUCN Red List of Threatened Species. 1964 - onwards. https://www.iucnredlist.org/ (Ref. 130435)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=137896&amp;speccode=4480'>Ford, M.0 &lt;i&gt;Lampetra fluviatilis&lt;/i&gt;. The IUCN Red List of Threatened Species 2024: e.T11206A135088436. (Ref. 137896)</reference><dataObject>
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	<agent homepage='http://www.fishbase.us/collaborators/CollaboratorSummary.php?id=7' logoURL='' role='photographer'>Robbie N. Cada</agent>
	<dcterms:created>1993-07-01</dcterms:created>
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	<agent homepage='' logoURL='' role='recorder'>Crispina B. Binohlan</agent>
	<dcterms:created>1991-06-21 00:00:00</dcterms:created>
	<dcterms:modified>2024-01-23 00:00:00</dcterms:modified>
	<dc:title>Biology</dc:title>
	<dc:language></dc:language>
	<license>http://creativecommons.org/licenses/by-nc/3.0/</license>
	<dc:rights></dc:rights>
	<dcterms:rightsHolder>WorldFish Center - FishBase</dcterms:rightsHolder>
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	<audience>General public</audience>
	<dc:source>http://www.fishbase.org/summary/SpeciesSummary.php?id=4480</dc:source>
	<subject>http://rs.tdwg.org/ontology/voc/SPMInfoItems#Migration</subject>
	<dc:description>In fresh waters, in rivers, brooks, and lakes.  Anadromous, but some populations are permanent freshwater residents (e.g. Lough Neagh, Northern Ireland (UK); Loch Lomond, Scotland (UK); lakes Ladoga and Onega, Russian Federation) (Ref. 89241).  Amphihaline species (Ref. 51442). Metamorphosis takes place at a length of about 13 cm and they migrate to the sea (Ref. 51442). Most river lampreys live 4-7 years and reach 30-35 cm length (Ref. 88187). Females grow larger than males (Ref. 58137).  Adults spend 1-2 years at sea, often along the coast or in estuaries (Ref. 59043). They live on hard bottoms or attached to larger fish like cod and herring (Ref. 88174).  Adults are parasitic, feeding on fishes by sucking their blood and afterwards consuming the flesh (Ref. 1998).  Adults predatory on marine fishes from the end of July to October.  Trematodes and cestodes have been found in the intestine of prespawning adults at sea. (Ref. 89241). In autumn, adults commonly undergo reproductive migration from the sea to shallow middle or upper reaches of rivers and streams with strong currents (1.0-2.0 m/s in British rivers) and gravel bottoms (Ref. 51442, 59043, 88171). Spawning migration upriver (Vistula and Neman rivers) begins near the end of September in the Baltic Sea Basin. In tributaries to the Gulf of Finland (Narva, Neva, and Luga rivers), upstream spawning migrations occur twice yearly; once in summer-fall and once in spring. In the upper Rhine River, the spawning season is February to April; in England (UK) from April to May; and in the lower Neva River, Russian Federation, from early June to early July (Ref. 89241).  During reproductive migration and reproduction, adults do not feed but instead utilize their lipid reserves (Ref. 30578) and are known to undergo a considerable shortening of up to 27 per cent (Ref. 83507).  Fecundity is highly variable and ranges from 650 to 42,500 eggs/female; 10,000-16,000 eggs/female in Lake Ladoga. Communal spawning in the same redd by L. fluviatilis and L. planeri has been reported in the River Tywi Basin, Wales (UK), in April, at a water temperature of 11 °C. Both species participated in constructing a redd about 23 cm in diameter and 5-8 cm deep, consisting of pebbles, gravel and coarse sand (Ref. 89241). Spawning takes place in pre-excavated pits in river beds, its depth between 50-100 cm; after spawning the adults die (Ref. 51442). The blind ammocoetes are filter feeders of detritus and microorganisms; they live mostly buried in sand, silt or clay sediments for up to 4.5 years, often at the edges of rivers and streams where currents are slow (Ref. 51442, 59043, 88184, 88185). Sometimes they are found in substrates with submerged vegetation and plant debris (Ref. 12285). Ammocoetes may tolerate low oxygen levels but when burrows reach near anoxic levels, the larvae need to emerge from the substrate in order to survive (Ref. 88184).  Fisheries exist in England (UK), Finland, France, and the Russian Federation. In the 19th century, up to 450,000 adults yearly were used by the English fishing fleet as bait in the fisheries for Gadus morhua and Psetta maxima. In Finland, the catch in 1983 was 2.3-2.4 million individuals (about 100 t) for a value of $800,000 US. There are reports of intoxication through eating this species (Halstead 1967). The mucus and serum are poisonous and the flesh must be thoroughly washed and all the blood removed before consumption (Ref. 5504). Cooking method involves de-sliming, removal of heads and branchial regions, the rest of the body covered in sunflower flour, and cooked in sunflower oil (Ref.89241). Utilized fresh and smoked; eaten fried (Ref. 9988).</dc:description>
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	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=3161&amp;speccode=4480'>Vladykov, V.D. 1984 Petromyzonidae. p. 64-67. In P.J.P. Whitehead, M.-L. Bauchot, J.-C. Hureau, J. Nielsen, and E. Tortonese (eds.) Fishes of the north-eastern Atlantic and Mediterranean. UNESCO, Paris. vol. 1. (Ref. 3161)</reference></dataObject><dataObject>
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	<agent homepage='http://oceans.ubc.ca/maria-lourdes-deng-palomares/' logoURL='' role='recorder'>Maria Lourdes D. Palomares</agent>
	<dcterms:created>1993-06-16 00:00:00</dcterms:created>
	<dcterms:modified>2013-10-31 00:00:00</dcterms:modified>
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	<dc:language></dc:language>
	<license>http://creativecommons.org/licenses/by-nc/3.0/</license>
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	<dcterms:rightsHolder>WorldFish Center - FishBase</dcterms:rightsHolder>
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	<audience>General public</audience>
	<dc:source>http://www.fishbase.org/summary/SpeciesSummary.php?id=4480</dc:source>
	<subject>http://rs.tdwg.org/ontology/voc/SPMInfoItems#TrophicStrategy</subject>
	<dc:description>It is jawless with a round sucker-like mouth and has an outer circle of small teeth and an inner circle of large teeth (Ref. 88171).  It has a typical eel-like shape with 2 dorsal fins and 7 gill openings behind the eye. It lacks paired fins. Young adults are uniformly greyish in colour. As it ages, the river lamprey becomes greenish-brown dorsally, golden yellow along the sides and white ventrally (Ref. 58137). In coastal waters of Germany, it can be confused with the sea lamprey (Petromyzon marinus), which is distinguished by having its teeth arranged in many consecutive circular rows (Ref. 88171). Other diagnostic features: Adults 8.6-49.2 cm TL. Body wet weight in individuals 18.0-49.2 cm TL, 30-150 g. Body proportions, as percentage of TL (based on 48 specimens measuring 10.8-38.6 cm TL): prebranchial length, 10.0-12.9; branchial length, 7.9-11.3; trunk length, 46.2-54.3; tail length, 24.1-30.3; eye length, 1.4-3.1; disc length, 4.6-7.0. Urogenital papilla length, as a percentage of branchial length, in 19 spawning males measuring 19.7-28.3 cm TL, 15.9-37.5. Trunk myomeres, 58-66. Dentition: marginals, 70-95; supraoral lamina, 2 unicuspid teeth; infraoral lamina, 5-9 either all unicuspid teeth or, more frequently, the lateralmost are bicuspid and the internal ones unicuspid; 3 endolaterals on each side; endolateral formula, typically 2-3-2, rarely 1-3-2 or 2-3-1; 1-2 rows of anterials; first row of anterials, 4-7 unicuspid teeth; exolaterals absent; posterials absent; transverse lingual lamina, 8-18, usually 12-14, unicuspid laminae straight or parentheses-shaped and each with 9-13 unicuspid teeth. Marginal membrane present. Velar tentacles, 4-10, with tubercles; no velar wings. Body coloration in recently metamorphosed individuals silvery; in preserved upstream migrants, bluish brown or lead gray on the dorsal aspect tending towards silvery on the lateral aspects and whitish or yellowish on ventral aspect. Early upstream spawning migrants returning from the sea have a bronze sheen. Dorsal fins of maturing individuals may have a purplish tint. Iris is golden yellow. Body coloration in the landlocked population in Lake Ladoga is completely black. Lateral line neuromasts unpigmented or darkly pigmented. Extent of caudal fin pigmentation, absent or trace in young adults and 75% or more in spawning individuals. Caudal fin shape, spade-like. Oral fimbriae, 84-112. Oral papillae, 11-20 (Ref. 89241).</dc:description>
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	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=2163&amp;speccode=4480'>Allardi, J. and P. Keith 1991 Atlas préliminaire des poissons d'eau douce de France. Coll. Patrimoines Naturels, vol. 4. Secrétariat Faune Flore, Muséum national d'Histoire naturelle, Paris. 234 p. (Ref. 2163)</reference></dataObject><dataObject>
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	<agent homepage='http://oceans.ubc.ca/maria-lourdes-deng-palomares/' logoURL='' role='recorder'>Maria Lourdes D. Palomares</agent>
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	<subject>http://rs.tdwg.org/ontology/voc/SPMInfoItems#Size</subject>
	<dc:description>Dorsal spines (total): 0;  Analspines: 0; Analsoft rays: 0</dc:description>
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	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=2163&amp;speccode=4480'>Allardi, J. and P. Keith 1991 Atlas préliminaire des poissons d'eau douce de France. Coll. Patrimoines Naturels, vol. 4. Secrétariat Faune Flore, Muséum national d'Histoire naturelle, Paris. 234 p. (Ref. 2163)</reference></dataObject><dataObject>
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	<agent homepage='' logoURL='' role='recorder'>Crispina B. Binohlan</agent>
	<dcterms:created>1991-06-21 00:00:00</dcterms:created>
	<dcterms:modified>2025-11-05 00:00:00</dcterms:modified>
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	<dc:language></dc:language>
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	<dcterms:rightsHolder>WorldFish Center - FishBase</dcterms:rightsHolder>
	<dcterms:bibliographicCitation></dcterms:bibliographicCitation>
	<audience>General public</audience>
	<dc:source>http://www.fishbase.org/summary/SpeciesSummary.php?id=4480</dc:source>
	<subject>http://rs.tdwg.org/ontology/voc/SPMInfoItems#Threats</subject>
	<dc:description>Europe: southern Norway to France, including Ireland and the British Isles. Also in the Baltic Sea and along the French and western Italian coasts of the Mediterranean Sea (Ref. 59043). Absent from Black, Caspian and Polar seas (Ref. 3161). Landlocked populations from Lake Mjosa in Norway (Ref. 12269), Lakes Ladoga and Onega, upper Volga in Russia, Loch Lomond in Scotland, some Finnish lakes and possibly in Lough Neagh in Ireland (Ref. 59043).</dc:description>
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	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=3161&amp;speccode=4480'>Vladykov, V.D. 1984 Petromyzonidae. p. 64-67. In P.J.P. Whitehead, M.-L. Bauchot, J.-C. Hureau, J. Nielsen, and E. Tortonese (eds.) Fishes of the north-eastern Atlantic and Mediterranean. UNESCO, Paris. vol. 1. (Ref. 3161)</reference></dataObject><dataObject>
	<dc:identifier>FB-Uses-4480</dc:identifier>
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	<agent homepage='' logoURL='' role='recorder'>Crispina B. Binohlan</agent>
	<dcterms:created>1991-06-21 00:00:00</dcterms:created>
	<dcterms:modified>2024-01-23 00:00:00</dcterms:modified>
	<dc:title>Importance</dc:title>
	<dc:language></dc:language>
	<license>http://creativecommons.org/licenses/by-nc/3.0/</license>
	<dc:rights></dc:rights>
	<dcterms:rightsHolder>WorldFish Center - FishBase</dcterms:rightsHolder>
	<dcterms:bibliographicCitation></dcterms:bibliographicCitation>
	<audience>General public</audience>
	<dc:source>http://www.fishbase.org/summary/SpeciesSummary.php?id=4480</dc:source>
	<subject>http://rs.tdwg.org/ontology/voc/SPMInfoItems#Diseases</subject>
	<dc:description>fisheries: minor commercial; bait: usually; price category: unknown; price reliability: </dc:description>
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	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=4645&amp;speccode=4480'>Muus, B. and P. Dahlström 1978 Meeresfische der Ostsee, der Nordsee, des Atlantiks. BLV Verlagsgesellschaft, München. 244 p. (Ref. 4645)</reference>
	<reference isbn='0-913235-53-9' url='http://www.fishbase.org/references/FBRefSummary.php?id=4537&amp;speccode=4480'>Robins, C.R., R.M. Bailey, C.E. Bond, J.R. Brooker, E.A. Lachner, R.N. Lea and W.B. Scott 1991 World fishes important to North Americans. Exclusive of species from the continental waters of the United States and Canada. Am. Fish. Soc. Spec. Publ. (21):243 p. (Ref. 4537)</reference></dataObject><dataObject>
	<dc:identifier>FB-Migration-4480</dc:identifier>
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	<agent homepage='' logoURL='' role='recorder'>Crispina B. Binohlan</agent>
	<dcterms:created>1991-06-21 00:00:00</dcterms:created>
	<dcterms:modified>2024-01-23 00:00:00</dcterms:modified>
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	<dcterms:rightsHolder>WorldFish Center - FishBase</dcterms:rightsHolder>
	<dcterms:bibliographicCitation></dcterms:bibliographicCitation>
	<audience>General public</audience>
	<dc:source>http://www.fishbase.org/summary/SpeciesSummary.php?id=4480</dc:source>
	<subject>http://rs.tdwg.org/ontology/voc/SPMInfoItems#Diseases</subject>
	<dc:description>Anadromous. Fish that ascend rivers to spawn, as salmon and hilsa do. Sub-division of diadromous.    Migrations should be cyclical and predictable and cover more than 100 km.</dc:description>
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	<thumbnailURL></thumbnailURL>
	<location></location>
	<reference isbn='978-2-8399-0298-4' url='http://www.fishbase.org/references/FBRefSummary.php?id=59043&amp;speccode=4480'>Kottelat, M. and J. Freyhof 2007 Handbook of European freshwater fishes. Publications Kottelat, Cornol and Freyhof, Berlin. 646 pp. (Ref. 59043)</reference></dataObject><dataObject>
	<dc:identifier>FB-LifeCycle-4480</dc:identifier>
	<dataType>http://purl.org/dc/dcmitype/Text</dataType>
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	<agent homepage='' logoURL='' role='recorder'>Sari Kuosmanen-Postila</agent>
	<dcterms:created>1991-06-21 00:00:00</dcterms:created>
	<dcterms:modified>2024-01-23 00:00:00</dcterms:modified>
	<dc:title></dc:title>
	<dc:language></dc:language>
	<license>http://creativecommons.org/licenses/by-nc/3.0/</license>
	<dc:rights></dc:rights>
	<dcterms:rightsHolder>WorldFish Center - FishBase</dcterms:rightsHolder>
	<dcterms:bibliographicCitation></dcterms:bibliographicCitation>
	<audience>General public</audience>
	<dc:source>http://www.fishbase.org//Reproduction/FishReproSummary.php?ID=4480&amp;StockCode=4678</dc:source>
	<subject>http://rs.tdwg.org/ontology/voc/SPMInfoItems#Diseases</subject>
	<dc:description>Adults migrate from the sea to the rivers to spawn (Ref. 51442), ascending rivers at night, anytime between autumn and spring but this movement ceases when water temperature drops (Ref. 59043). During the early stages of its spawning migration, the river lamprey has been shown to be attracted to chemical odours given off by ammocoetes (Ref. 88188). Males reach the spawning grounds first and build nests at depths between 50-100 cm, which have a diameter of 20-40 cm and a height of 10 cm (Ref. 12285, 51442, 59043, 88186). The river lamprey forms spawning aggregations, often during sunny days, when water temperature rises above 9 °C (Ref. 59043). Females may spawn with up to 6 males on separate occasions. One nest is usually utilized by a single mating pair (Ref. 59043). Adults spawn only once in their lifetime and usually die within 2 weeks after spawning (Ref. 12285, 51442). Eggs hatch in 15-30 days (Ref. 88184). The ammocoetes live in the calm zones of the river until metamorphosis at about 13 cm length before migrating to the sea (Ref. 51442).</dc:description>
	<mediaURL></mediaURL>
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	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=6114&amp;speccode=4480'>Koli, L. 1990 Suomen kalat. [Fishes of Finland]. Werner Söderström Osakeyhtiö. Helsinki. 357 p. (in Finnish). (Ref. 6114)</reference></dataObject><dataObject>
	<dc:identifier>FB-TrophicStrategy-4480</dc:identifier>
	<dataType>http://purl.org/dc/dcmitype/Text</dataType>
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	<agent homepage='' logoURL='' role='recorder'>Susan M. Luna</agent>
	<dcterms:created>1991-10-18 00:00:00</dcterms:created>
	<dcterms:modified>2013-10-31 00:00:00</dcterms:modified>
	<dc:title></dc:title>
	<dc:language></dc:language>
	<license>http://creativecommons.org/licenses/by-nc/3.0/</license>
	<dc:rights></dc:rights>
	<dcterms:rightsHolder>WorldFish Center - FishBase</dcterms:rightsHolder>
	<dcterms:bibliographicCitation></dcterms:bibliographicCitation>
	<audience>General public</audience>
	<dc:source>fluviatilis</dc:source>
	<subject>http://rs.tdwg.org/ontology/voc/SPMInfoItems#Diseases</subject>
	<dc:description>Adults are hematophageous predators (Ref. 51442).  They often inflict extensive damage on their hosts by rasping away large amounts of flesh from the back; they become bloated when feeding because the entire gut is full of blood and fish flesh (Ref. 1998).  The ammocoetes larvae are microphageous and feed on diatoms and protozoans (Ref. 51442).</dc:description>
	<mediaURL></mediaURL>
	<thumbnailURL></thumbnailURL>
	<location></location>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=682&amp;speccode=4480'>Muus, B.J. and P. Dahlström 1967 Guide des poissons d'eau douce et pêche. GEC Gads Forlag, Copenhague. 242 p. (Ref. 682)</reference></dataObject><dataObject>
	<dc:identifier>FB-Size-4480</dc:identifier>
	<dataType>http://purl.org/dc/dcmitype/Text</dataType>
	<mimeType>text/html</mimeType>
	<agent homepage='' logoURL='' role='recorder'>Crispina B. Binohlan</agent>
	<dcterms:created>1991-06-21 00:00:00</dcterms:created>
	<dcterms:modified>2024-01-23 00:00:00</dcterms:modified>
	<dc:title>Max. size</dc:title>
	<dc:language></dc:language>
	<license>http://creativecommons.org/licenses/by-nc/3.0/</license>
	<dc:rights></dc:rights>
	<dcterms:rightsHolder>WorldFish Center - FishBase</dcterms:rightsHolder>
	<dcterms:bibliographicCitation></dcterms:bibliographicCitation>
	<audience>General public</audience>
	<dc:source>http://www.fishbase.org/summary/SpeciesSummary.php?id=4480</dc:source>
	<subject>http://rs.tdwg.org/ontology/voc/SPMInfoItems#Diseases</subject>
	<dc:description>50.0 cm TL (male/unsexed; (Ref. 30578)); max. published weight: 150 g (Ref. 40476); max. reported age: 10 years (Ref. 30578)</dc:description>
	<mediaURL></mediaURL>
	<thumbnailURL></thumbnailURL>
	<location></location>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=30578&amp;speccode=4480'>Billard, R. 1997 Les poissons d'eau douce des rivières de France. Identification, inventaire et répartition des 83 espèces. Lausanne, Delachaux &amp; Niestlé, 192p. (Ref. 30578)</reference>
	<reference isbn='2-85653653261' url='http://www.fishbase.org/references/FBRefSummary.php?id=40476&amp;speccode=4480'>Keith, P. and J. Allardi (coords.) 2001 Atlas des poissons d'eau douce de France. Muséum national d’Histoire naturelle, Paris. Patrimoines naturels, 47:1-387. (Ref. 40476)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=30578&amp;speccode=4480'>Billard, R. 1997 Les poissons d'eau douce des rivières de France. Identification, inventaire et répartition des 83 espèces. Lausanne, Delachaux &amp; Niestlé, 192p. (Ref. 30578)</reference></dataObject><dataObject>
	<dc:identifier>FB-Threats-2</dc:identifier>
	<dataType>http://purl.org/dc/dcmitype/Text</dataType>
	<mimeType>text/html</mimeType>
	<agent homepage='' logoURL='' role='recorder'>Belen Acosta</agent>
	<dcterms:created>1990-10-18 00:00:00</dcterms:created>
	<dcterms:modified>2024-03-07 00:00:00</dcterms:modified>
	<dc:title></dc:title>
	<dc:language></dc:language>
	<license>http://creativecommons.org/licenses/by-nc/3.0/</license>
	<dc:rights></dc:rights>
	<dcterms:rightsHolder>WorldFish Center - FishBase</dcterms:rightsHolder>
	<dcterms:bibliographicCitation></dcterms:bibliographicCitation>
	<audience>General public</audience>
	<dc:source>http://www.fishbase.org/summary/SpeciesSummary.php?id=2</dc:source>
	<subject>http://rs.tdwg.org/ontology/voc/SPMInfoItems#Diseases</subject>
	<dc:description>Least Concern (LC) , IUCN Grouper and Wrasse Specialist Group</dc:description>
	<mediaURL></mediaURL>
	<thumbnailURL></thumbnailURL>
	<location></location>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=57073'>IUCN 2006 2006 IUCN red list of threatened species. www.iucnredlist.org. Downloaded July 2006.</reference></dataObject><dataObject>
	<dc:identifier>FB-Habitat-4480</dc:identifier>
	<dataType>http://purl.org/dc/dcmitype/Text</dataType>
	<mimeType>text/html</mimeType>
	<agent homepage='' logoURL='' role='recorder'>Crispina B. Binohlan</agent>
	<dcterms:created>1991-06-21 00:00:00</dcterms:created>
	<dcterms:modified>2024-01-23 00:00:00</dcterms:modified>
	<dc:title>Environment</dc:title>
	<dc:language></dc:language>
	<license>http://creativecommons.org/licenses/by-nc/3.0/</license>
	<dc:rights></dc:rights>
	<dcterms:rightsHolder>WorldFish Center - FishBase</dcterms:rightsHolder>
	<dcterms:bibliographicCitation></dcterms:bibliographicCitation>
	<audience>General public</audience>
	<dc:source>http://www.fishbase.org/summary/SpeciesSummary.php?id=4480</dc:source>
	<subject>http://rs.tdwg.org/ontology/voc/SPMInfoItems#Habitat</subject>
	<dc:description>demersal; anadromous (Ref. 59043); freshwater; brackish; marine; depth range 10 - ? m</dc:description>
	<mediaURL></mediaURL>
	<thumbnailURL></thumbnailURL>
	<location></location>
	<reference isbn='978-2-8399-0298-4' url='http://www.fishbase.org/references/FBRefSummary.php?id=59043&amp;speccode=4480'>Kottelat, M. and J. Freyhof 2007 Handbook of European freshwater fishes. Publications Kottelat, Cornol and Freyhof, Berlin. 646 pp. (Ref. 59043)</reference></dataObject>
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