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	<dc:identifier>FB-3445</dc:identifier>
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	<dwc:Kingdom>Animalia</dwc:Kingdom>
	<dwc:Phylum>Chordata</dwc:Phylum>
	<dwc:Class>Teleostei</dwc:Class>
	<dwc:Order>Perciformes/Percoidei</dwc:Order>
	<dwc:Family>Percidae</dwc:Family>
	<dwc:Genus>Etheostoma</dwc:Genus>
	<dwc:ScientificName>Etheostoma nigrum Rafinesque, 1820</dwc:ScientificName>
	<commonName xml:lang='da'>Johnny-smutte</commonName>
	<commonName xml:lang='de'>Schwarzer Springbarsch</commonName>
	<commonName xml:lang='en'>Johnny darter</commonName>
	<commonName xml:lang='et'>Johnny darter</commonName>
	<commonName xml:lang='et'>Mustjas pärisdarter</commonName>
	<commonName xml:lang='fr'>Raseux-de-terre noir</commonName>
	<commonName xml:lang=''>黑鏢鱸</commonName>
	<commonName xml:lang=''>黑镖鲈</commonName>
	<synonym relationship='synonym'>Boleosoma nigrum (Rafinesque, 1820)</synonym>
	<synonym relationship='synonym'>Boleosoma nigrum nigrum (Rafinesque, 1820)</synonym>
	<synonym relationship='synonym'>Etheostoma nigra Rafinesque, 1820</synonym>
	<synonym relationship='synonym'>Etheostoma nigrum nigrum Rafinesque, 1820</synonym>
	<dcterms:created>1991-05-01</dcterms:created>
	<dcterms:modified>2008-12-29</dcterms:modified>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=276&amp;speccode=3445'>Robins, C.R., R.M. Bailey, C.E. Bond, J.R. Brooker, E.A. Lachner, R.N. Lea and W.B. Scott0 A list of common and scientific names of fishes from the United States and Canada. Am. Fish. Soc. Spec. Publ. (12)1-174. (Ref. 276)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=1998&amp;speccode=3445'>Scott, W.B. and E.J. Crossman0 Freshwater fishes of Canada. Bull. Fish. Res. Board Can. 184: xi+1-966. (Ref. 1998)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=2059&amp;speccode=3445'>Baensch, H.A. and R. Riehl0 Aquarien atlas. Bd. 3. Melle: Mergus, Verlag für Natur-und Heimtierkunde, Germany. 1104 p. (Ref. 2059)</reference>
	<reference isbn='0-913235-69-5 (pbk).' url='http://www.fishbase.org/references/FBRefSummary.php?id=3814&amp;speccode=3445'>Robins, C.R., R.M. Bailey, C.E. Bond, J.R. Brooker, E.A. Lachner, R.N. Lea and W.B. Scott0 Common and scientific names of fishes from the United States and Canada. Am. Fish. Soc. Spec. Publ. (20):183 p. (Ref. 3814)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=4517&amp;speccode=3445'>Hureau, J.-C.0 La base de données GICIM: Gestion informatisée des collections ichthyologiques du Muséum. p. 225-227. In Atlas Préliminaire des Poissons d'Eau Douce de France. Conseil Supérieur de la Pêche, Ministère de l'Environnement, CEMAGREF et Muséum national d'Histoire naturelle, Paris. (Ref. 4517)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=5723&amp;speccode=3445'>Page, L.M. and B.M. Burr0 A field guide to freshwater fishes of North America north of Mexico. Houghton Mifflin Company, Boston. 432 p. (Ref. 5723)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=7043&amp;speccode=3445'>Page, L.0 Handbook of darters. T.F.H. Publications, Inc. USA. 271 p. (Ref. 7043)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=8973&amp;speccode=3445'>Vasil'ev, V.P.0 Chromosome numbers in fish-like vertebrates and fish. J. Ichthyol. 20(3):1-38. (Ref. 8973)</reference>
	<reference isbn='0-87049-711-1' url='http://www.fishbase.org/references/FBRefSummary.php?id=10294&amp;speccode=3445'>Etnier, D.A. and W.C. Starnes0 The fishes of Tennessee. The University of Tennessee Press, Knoxville, Tennessee, USA. (pls. check date). (Ref. 10294)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=10937&amp;speccode=3445'>Anonymous0 Fish collection database of the University of British Columbia Fish Museum Fish Museum. University of British Columbia, Vancouver, Canada. (Ref. 10937)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=12193&amp;speccode=3445'>Hugg, D.O.0 MAPFISH georeferenced mapping database. Freshwater and estuarine fishes of North America. Life Science Software. Dennis O. and Steven Hugg, 1278 Turkey Point Road, Edgewater, Maryland, USA. (Ref. 12193)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=12204&amp;speccode=3445'>Coad, B.W.0 Encyclopedia of Canadian fishes. Canadian Museum of Nature and Canadian Sportfishing Productions Inc. Singapore. (Ref. 12204)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=26282&amp;speccode=3445'>Eschmeyer, W.N. (ed.)0 Catalog of fishes. Special Publication, California Academy of Sciences, San Francisco. 3 vols. 2905 p. (Ref. 26282)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=29212&amp;speccode=3445'>Ross, M.R.0 A chromosome study of five species of Etheostominae fishes (Percidae). Copeia 1973(1):163-165. (Ref. 29212)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=30184&amp;speccode=3445'>Arkhipchuk, V.V.0 Chromosome database. Database of Dr. Victor Arkhipchuk. (Ref. 30184)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=30611&amp;speccode=3445'>Facey, D.E. and G.W. Labar0 Biology of American eels in Lake Champlain, Vermont. Trans. Am. Fish. Soc. 110(3):396-402. (Ref. 30611)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=30612&amp;speccode=3445'>Jude, D.J., F.J. Tesar, S.F. Deboe and T.J. Miller0 Diet and selection of major prey species by Lake Michigan salmonids, 1973-1982. Trans. Am. Fish. Soc. 116(5):677-691. (Ref. 30612)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=31982&amp;speccode=3445'>Anonymous0 Fish collection database of the Natural History Museum, London (formerly British Museum of Natural History (BMNH)). Natural History Museum, London (formerly British Museum of Natural History (BMNH)). (Ref. 31982)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=34370&amp;speccode=3445'>Klinkhardt, M, M. Tesche and H. Greven0 Database of fish chromosomes. Westarp Wissenschaften. (Ref. 34370)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=34541&amp;speccode=3445'>Danzmann, R.G.0 The karyology of eight species of fish belonging to the family Percidae. Can. J. Zool. 57:2055-2060. (Ref. 34541)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=34634&amp;speccode=3445'>Anonymous0 Fish collection database of the Gulf Coast Research Laboratory (GCRL). The Gulf Coast Research Laboratory (GCRL), Ocean Springs, Mississippi, USA. (Ref. 34634)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=35064&amp;speccode=3445'>Ross, S.T.0 The systematics of &lt;i&gt;Gasterosteus aculeatus&lt;/i&gt; (Pisces, Gasterosteidae) in central and southern Califormia. Natur. Hist. Mus. Los Angeles Country. Contrib. Sci. 243:1-20. (Ref. 35064)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=35508&amp;speccode=3445'>Anonymous0 The icthyological collection of the Zoological Museum Hamburg (ZMH). Division of Ichthyology and Herpetology, Zoological Museum Hamburg (ZMH). (Ref. 35508)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=36670&amp;speccode=3445'>Anonymous0 Fish collection database of the J.L.B. Smith Institute of Ichthyology, Grahamstown, South Africa. J.L.B. Smith Institute of Ichthyology, Grahamstown, South Africa. (Ref. 36670)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=37092&amp;speccode=3445'>Lotrich, V.A.0 Growth, production, and community composition of fishes inhabiting a first-, second-, and third-order stream of eastern Kentucky. Ecol. Monogr. 43:377-397. (Ref. 37092)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=38712&amp;speccode=3445'>Martin, F.D.0 Diets of four sympatric species of &lt;i&gt;Etheostoma&lt;/i&gt; (Pisces: Percidae) from southern Indiana: interspecific and intraspecific multiple comparisons. Environ. Biol. Fishes 11(2):113-120. (Ref. 38712)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=38732&amp;speccode=3445'>Anonymous0 Fish collection database of the National Museum of Natural History (Smithsonian Institution). Smithsonian Institution-Division of Fishes. (Ref. 38732)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=41414&amp;speccode=3445'>Anonymous0 Fish collection database of the American Museum of Natural History. American Museum of Natural History, Central Park West , NY 10024-5192, USA. (Ref. 41414)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=42125&amp;speccode=3445'>Anonymous0 Fish collection of the Royal British Columbia Museum. Royal British Columbia Museum, 675 Belleville Street, Victoria BC, Canada V8V 1X4. (Ref. 42125)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=46644&amp;speccode=3445'>Miller, J.E., J.F. Savino and R.K. Neely0 Competition for food between crayfish (&lt;i&gt;Orconectes virilis&lt;/i&gt;) and the slimy sculpin (&lt;i&gt;Cottus cognatus&lt;/i&gt;). J. Freshwat. Ecol. 7(2):127-135. (Ref. 46644)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=47438&amp;speccode=3445'>Anonymous0 Fish collection of the Royal Ontario Museum. Royal Ontario Museum. (Ref. 47438)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=50685&amp;speccode=3445'>Flemer, D.A. and W.S. Woolcott0 Food habits and distribution of the fishes of Tuckahoe Creek, Virginia, with special emphasis on the bluegill, &lt;i&gt;Lepomis m. macrochirus&lt;/i&gt; Rafinesque. Chesapeake Sci. 7(2):75-89. (Ref. 50685)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=51471&amp;speccode=3445'>Carl, H.0 Danish fish names. Zoological Museum of Copenhagen. Unpublished. (Ref. 51471)</reference>
	<reference isbn='1-888569-61-1' url='http://www.fishbase.org/references/FBRefSummary.php?id=52299&amp;speccode=3445'>Nelson, J.S., E.J. Crossman, H. Espinosa-Pérez, L.T. Findley, C.R. Gilbert, R.N. Lea and J.D. Williams0 Common and scientific names of fishes from the United States, Canada, and Mexico. American Fisheries Society, Special Publication 29, Bethesda, Maryland. ix, 386 p. + 1 CD. (Ref. 52299)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=52559&amp;speccode=3445'>Coker, G.A., C.B. Portt and C.K. Minns0 Morphological and ecological characteristics of Canadian freshwater fishes. Can. MS Rpt. Fish. Aquat. Sci. 2554:iv+89p. (Ref. 52559)</reference>
	<reference isbn='0-299-08790-5' url='http://www.fishbase.org/references/FBRefSummary.php?id=54725&amp;speccode=3445'>Becker, G.C.0 Fishes of Wisconsin. Madison (USA, WI): University of Wisconsin Press, xii, 1052 p. (Ref. 54725)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=58108&amp;speccode=3445'>Chinese Academy of Fishery Sciences0 Chinese aquatic germplasm resources database. http://zzzy.fishinfo.cn (Ref. 58108)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=74952&amp;speccode=3445'>Winn, H.E.0 Comparative reproductive behavior and ecology of fourteen species of darters (Pisces-Percidae). Ecol. Monogr. 28:155-191. (Ref. 74952)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=74974&amp;speccode=3445'>Smith, P.W. and L.M. Page0 The food of spotted bass in streams of the Wabash River drainage. American Fisheries Society of Transactions 98(4):647-651. (Ref. 74974)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=74979&amp;speccode=3445'>Lutterbie, G.W.0 Reproduction and age and growth in Wisconsin darters (Osteichthyes: Percidae). Rep. Fauna Flora Wis. 15. 44 p. (Ref. 74979)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=74980&amp;speccode=3445'>Speare, E.P.0 Fecundity and egg survival of the central Johnny darter (&lt;i&gt;Etheostoma nigrum nigrum&lt;/i&gt;) in southern Michigan. Copeia 1965(3):308-314. (Ref. 74980)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=74981&amp;speccode=3445'>Raney, E.C. and E.A. Lachner0 Observations on the life history of the spotted darter, &lt;i&gt;Poecilichthys maculatus&lt;/i&gt; (Kirtland). Copeia 1939(3):157-165. (Ref. 74981)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=78789&amp;speccode=3445'>Missouri Department of Conservation0 Fish of Missouri. Missouri Department of Conservation. (Ref. 78789)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=105471&amp;speccode=3445'>eElurikkus0 eElurikkus koondab ühte portaali Eesti eluslooduse andmed. http://elurikkus.ut.ee/ (Ref. 105471)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=105574&amp;speccode=3445'>Termeki on Eesti Keele Instituudi0 Termeki on Eesti Keele Instituudi. http://term.eki.ee/termbase/view/7351963/ (Ref. 105574)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=131016&amp;speccode=3445'>Driehaus, E.R., J. Landreth, K. Adase, D. Smith, D. Wellman, C.C. Arantes and B.A. Murry0 Length-weight relationship for 44 Central Appalachian fish species. J. Appl. Ichthyol. (Ref. 131016)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=132215&amp;speccode=3445'>WDNR, Wisconsin Department of Natural Resources0 Wisconsin Fish Species List.. World Wide Web electronic publication, Accessed 05 Oct. 2024. Madison (USA, WI):  Wisconsin Department of Natural Resources, 6 p. https://dnr.wisconsin.gov/sites/default/files/topic/Fishing/Species_wifish.pdf (Ref. 132215)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=137577&amp;speccode=3445'>Price, S.A., S.T. Friedman, K.A. Corn, O. Larouche, K. Brockelsby, A.J. Lee, M. Nagaraj, N.G. Bertrand, M. Danao, M.C. Coyne, J.R. Estrada, R. Friedman, E. Hoeft, M. Iwan, D. Gross, J.H. Kao, B. Landry, M.J. Linares, C. McGlinn and …0 FishShapes v1: Functionally relevant measurements of teleost shape and size on three dimensions. Ecology 103(e3829):1-2. (Ref. 137577)</reference><dataObject>
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	<dc:description>Occur in sandy and muddy, sometimes rocky, pools of headwaters, creeks, and small to medium rivers; and in sandy shores of lakes (Ref. 5723, 10294); also found in streams (Ref. 10294).  Adults feed on midge larvae, mayfly nymphs, caddis larvae, and microcrustaceans; young on entomostracans and tiny midge larvae (Ref. 10294).  Eggs are found clustered on underside of stone and guarded by males (Ref. 7043).</dc:description>
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	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=5723&amp;speccode=3445'>Page, L.M. and B.M. Burr 1991 A field guide to freshwater fishes of North America north of Mexico. Houghton Mifflin Company, Boston. 432 p. (Ref. 5723)</reference></dataObject><dataObject>
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	<dc:description>North America:  St. Lawrence-Great Lakes, Hudson Bay, and Mississippi River basins from Hudson Bay to southern Mississippi in the USA, and from Quebec in Canada and from Virginia in the USA to Saskatchewan in Canada and Colorado in the USA; and on Atlantic and Gulf Slope drainages in the USA.</dc:description>
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	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=5723&amp;speccode=3445'>Page, L.M. and B.M. Burr 1991 A field guide to freshwater fishes of North America north of Mexico. Houghton Mifflin Company, Boston. 432 p. (Ref. 5723)</reference></dataObject><dataObject>
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	<dc:description>Eggs are found clustered on underside of stone (Ref. 7043).  Males guard the eggs (Ref. 7043).</dc:description>
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	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=7043&amp;speccode=3445'>Page, L. 1983 Handbook of darters. T.F.H. Publications, Inc. USA. 271 p. (Ref. 7043)</reference></dataObject><dataObject>
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	<dc:description>Occurs in sandy and muddy, sometimes rocky, pools of headwaters, creeks, and small to medium rivers; and in sandy shores of lakes (Ref. 5723).</dc:description>
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	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=38712&amp;speccode=3445'>Martin, F.D. 1984 Diets of four sympatric species of &lt;i&gt;Etheostoma&lt;/i&gt; (Pisces: Percidae) from southern Indiana: interspecific and intraspecific multiple comparisons. Environ. Biol. Fishes 11(2):113-120. (Ref. 38712)</reference></dataObject><dataObject>
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	<dc:title>Max. size</dc:title>
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	<dc:description>7.2 cm TL (male/unsexed; (Ref. 5723)); max. reported age: 4 years (Ref. 12193)</dc:description>
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	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=5723&amp;speccode=3445'>Page, L.M. and B.M. Burr 1991 A field guide to freshwater fishes of North America north of Mexico. Houghton Mifflin Company, Boston. 432 p. (Ref. 5723)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=12193&amp;speccode=3445'>Hugg, D.O. 1996 MAPFISH georeferenced mapping database. Freshwater and estuarine fishes of North America. Life Science Software. Dennis O. and Steven Hugg, 1278 Turkey Point Road, Edgewater, Maryland, USA. (Ref. 12193)</reference></dataObject><dataObject>
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	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=57073'>IUCN 2006 2006 IUCN red list of threatened species. www.iucnredlist.org. Downloaded July 2006.</reference></dataObject><dataObject>
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	<dc:source>http://www.fishbase.org/summary/SpeciesSummary.php?id=3445</dc:source>
	<subject>http://rs.tdwg.org/ontology/voc/SPMInfoItems#Threats</subject>
	<dc:description>benthopelagic; freshwater; pH range: 7.0; dH range: 15</dc:description>
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