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<taxon>
	<dc:identifier>FB-2059</dc:identifier>
	<dc:source>http://www.fishbase.org/summary/SpeciesSummary.php?id=2059</dc:source>
	<dwc:Kingdom>Animalia</dwc:Kingdom>
	<dwc:Phylum>Chordata</dwc:Phylum>
	<dwc:Class>Elasmobranchii</dwc:Class>
	<dwc:Order>Rajiformes</dwc:Order>
	<dwc:Family>Rajidae</dwc:Family>
	<dwc:Genus>Raja</dwc:Genus>
	<dwc:ScientificName>Raja clavata Linnaeus, 1758</dwc:ScientificName>
	<commonName xml:lang='no'>Piggrokke</commonName>
	<commonName xml:lang='no'>Piggskate</commonName>
	<commonName xml:lang='pl'>Płaszczka nabijana</commonName>
	<commonName xml:lang='pl'>Raja ciernista</commonName>
	<commonName xml:lang='pl'>Raja nabijana</commonName>
	<commonName xml:lang='pt'>Arraia</commonName>
	<commonName xml:lang='pt'>Lenga</commonName>
	<commonName xml:lang='pt'>Raia-lenga</commonName>
	<commonName xml:lang='pt'>Raia</commonName>
	<commonName xml:lang='pt'>Raia lenga</commonName>
	<commonName xml:lang='pt'>Raia-brocheada</commonName>
	<commonName xml:lang='pt'>Raia-pinta</commonName>
	<commonName xml:lang='af'>Doringrug-rog</commonName>
	<commonName xml:lang='af'>Rog</commonName>
	<commonName xml:lang='sq'>Dhelpra e detit</commonName>
	<commonName xml:lang='sq'>Ragja</commonName>
	<commonName xml:lang='sq'>Raja</commonName>
	<commonName xml:lang='sq'>Raja guresh</commonName>
	<commonName xml:lang='sq'>Shori</commonName>
	<commonName xml:lang='ar'>Bakra</commonName>
	<commonName xml:lang='ar'>Baqara Msaffah'a</commonName>
	<commonName xml:lang='co'>Razza spinosa</commonName>
	<commonName xml:lang='cy'>Morgath ttyds</commonName>
	<commonName xml:lang='da'>Sømrokke</commonName>
	<commonName xml:lang='de'>Dornrochen</commonName>
	<commonName xml:lang='de'>Keulenrochen</commonName>
	<commonName xml:lang='de'>Keulenstachel</commonName>
	<commonName xml:lang='de'>Keulen-Stachel</commonName>
	<commonName xml:lang='de'>Nagelroche</commonName>
	<commonName xml:lang='de'>Nagelrochen</commonName>
	<commonName xml:lang='de'>Stachelrochen</commonName>
	<commonName xml:lang='de'>Steinrochen</commonName>
	<commonName xml:lang='en'>Maiden ray</commonName>
	<commonName xml:lang='en'>Rocker</commonName>
	<commonName xml:lang='en'>Roker</commonName>
	<commonName xml:lang='en'>Skate</commonName>
	<commonName xml:lang='en'>Thorny</commonName>
	<commonName xml:lang='en'>Thornback</commonName>
	<commonName xml:lang='en'>Thornback maid ray</commonName>
	<commonName xml:lang='en'>Thornback ray</commonName>
	<commonName xml:lang='en'>Thornback rough ray</commonName>
	<commonName xml:lang='en'>Thornback skate</commonName>
	<commonName xml:lang='et'>Ogarai</commonName>
	<commonName xml:lang='fo'>Naglaskøta</commonName>
	<commonName xml:lang='fi'>Okarausku</commonName>
	<commonName xml:lang='fr'>Clavelade</commonName>
	<commonName xml:lang='fr'>Raie bouclée</commonName>
	<commonName xml:lang='fr'>Raie bouclée clovée</commonName>
	<commonName xml:lang='fr'>Raiton</commonName>
	<commonName xml:lang='fr'>Réagris</commonName>
	<commonName xml:lang='fr'>Zerra bouké</commonName>
	<commonName xml:lang='hr'>Barakula</commonName>
	<commonName xml:lang='hr'>Kamenica</commonName>
	<commonName xml:lang='hr'>Klinka</commonName>
	<commonName xml:lang='hr'>Polig</commonName>
	<commonName xml:lang='hr'>Raza</commonName>
	<commonName xml:lang='hr'>Raža</commonName>
	<commonName xml:lang='hr'>Raža kamenica</commonName>
	<commonName xml:lang='is'>Dröfnuskata</commonName>
	<commonName xml:lang='it'>Arzilla di scoglio</commonName>
	<commonName xml:lang='it'>Arzilla pietrosa</commonName>
	<commonName xml:lang='it'>Baracola</commonName>
	<commonName xml:lang='it'>Baracoleta</commonName>
	<commonName xml:lang='it'>Perdosa</commonName>
	<commonName xml:lang='it'>Picara pitrusa</commonName>
	<commonName xml:lang='it'>Picareia</commonName>
	<commonName xml:lang='it'>Pichira pitrusa</commonName>
	<commonName xml:lang='it'>Raggia</commonName>
	<commonName xml:lang='it'>Ragia pitrusa</commonName>
	<commonName xml:lang='it'>Raia petrosa</commonName>
	<commonName xml:lang='it'>Raja petrusa</commonName>
	<commonName xml:lang='it'>Raja pitrusa</commonName>
	<commonName xml:lang='it'>Rasa</commonName>
	<commonName xml:lang='it'>Rasa spinosa</commonName>
	<commonName xml:lang='it'>Rasca petrusa</commonName>
	<commonName xml:lang='it'>Rascia petroèse</commonName>
	<commonName xml:lang='it'>Raza</commonName>
	<commonName xml:lang='it'>Raza pietrosa</commonName>
	<commonName xml:lang='it'>Razza</commonName>
	<commonName xml:lang='it'>Razza chiodata</commonName>
	<commonName xml:lang='it'>Razza di scoglio</commonName>
	<commonName xml:lang='it'>Razza spinusa</commonName>
	<commonName xml:lang='it'>Scardano</commonName>
	<commonName xml:lang='it'>Scritta perdosa</commonName>
	<commonName xml:lang='mg'>Fay</commonName>
	<commonName xml:lang='mg'>Makoba</commonName>
	<commonName xml:lang='mt'>Raja petruza</commonName>
	<commonName xml:lang='mt'>Raja tal-fosos</commonName>
	<commonName xml:lang='ru'>Morskaya lisitsa</commonName>
	<commonName xml:lang='ru'>лисица морская</commonName>
	<commonName xml:lang='ru'>Лисица морская (=скат колючий)</commonName>
	<commonName xml:lang='ru'>скат шиповатый</commonName>
	<commonName xml:lang='sr'>Raza kamenjarka</commonName>
	<commonName xml:lang=''>Dinnan</commonName>
	<commonName xml:lang=''>Doctot</commonName>
	<commonName xml:lang=''>Flair</commonName>
	<commonName xml:lang=''>Gunnack</commonName>
	<commonName xml:lang=''>Gurney</commonName>
	<commonName xml:lang=''>Thornie</commonName>
	<commonName xml:lang=''>Wildmeer skate</commonName>
	<commonName xml:lang='sv'>Knaggrocka</commonName>
	<commonName xml:lang='tr'>Deniztilkisi balığı</commonName>
	<commonName xml:lang='tr'>Dikenli vatoz</commonName>
	<commonName xml:lang='tr'>İğnelivatoz balığı</commonName>
	<commonName xml:lang='tr'>Vatoz</commonName>
	<commonName xml:lang='tr'>Vatoz balığı</commonName>
	<commonName xml:lang='uk'>Morska lysitsa</commonName>
	<commonName xml:lang='uk'>Морська лисиця</commonName>
	<commonName xml:lang='uk'>Плахур шипуватий</commonName>
	<commonName xml:lang='uk'>Скат шипуватий</commonName>
	<commonName xml:lang='wo'>Thiaukher</commonName>
	<commonName xml:lang='es'>Arraizabala</commonName>
	<commonName xml:lang='es'>Bastina rajada</commonName>
	<commonName xml:lang='es'>Bramante</commonName>
	<commonName xml:lang='es'>Clavel</commonName>
	<commonName xml:lang='es'>Gastaka</commonName>
	<commonName xml:lang='es'>Pez de mahoma</commonName>
	<commonName xml:lang='es'>Raia de clavos</commonName>
	<commonName xml:lang='es'>Rajada</commonName>
	<commonName xml:lang='es'>Rajada vera</commonName>
	<commonName xml:lang='es'>Raya común</commonName>
	<commonName xml:lang='es'>Raya crabuda</commonName>
	<commonName xml:lang='es'>Raya de clavos</commonName>
	<commonName xml:lang='es'>Raya punjosa</commonName>
	<commonName xml:lang='es'>Romaguera</commonName>
	<commonName xml:lang=''>Clavellada</commonName>
	<commonName xml:lang=''>Kalkanovatos</commonName>
	<commonName xml:lang=''>Raïa</commonName>
	<commonName xml:lang=''>Raža trnjevka</commonName>
	<commonName xml:lang=''>Sálahi</commonName>
	<commonName xml:lang=''>Stekelrog</commonName>
	<commonName xml:lang=''>Vatos</commonName>
	<commonName xml:lang=''>Vátos</commonName>
	<commonName xml:lang=''>Vulpe de mare</commonName>
	<commonName xml:lang=''>Βάτος</commonName>
	<commonName xml:lang=''>Καλκανόβατος</commonName>
	<commonName xml:lang=''>Καρκαβάκα</commonName>
	<commonName xml:lang=''>Ράγια</commonName>
	<commonName xml:lang=''>Ράσα</commonName>
	<commonName xml:lang=''>Ράτσα</commonName>
	<commonName xml:lang=''>Ρίνα</commonName>
	<commonName xml:lang=''>Σαλάχι</commonName>
	<commonName xml:lang=''>背棘鰩</commonName>
	<commonName xml:lang=''>背棘鳐</commonName>
	<synonym relationship='synonym'>Cephaleutherus maculatus Rafinesque, 1810</synonym>
	<synonym relationship='misapplied name'>Dasybatis asterias (non Delaroche, 1809)</synonym>
	<synonym relationship='synonym'>Hieroptera abredonensis Fleming, 1841</synonym>
	<synonym relationship='synonym'>Malacoraja clavata (Linnaeus, 1758)</synonym>
	<synonym relationship='synonym'>Raia aspera Risso, 1810</synonym>
	<synonym relationship='misapplied name'>Raia barnardi (non Norman, 1935)</synonym>
	<synonym relationship='synonym'>Raia clavata Linnaeus, 1758</synonym>
	<synonym relationship='synonym'>Raia rhizacanthus Regan, 1906</synonym>
	<synonym relationship='synonym'>Raia rubus Bloch, 1784</synonym>
	<synonym relationship='synonym'>Raja aspera Risso, 1810</synonym>
	<synonym relationship='misapplied name'>Raja asterias (non Delaroche, 1809)</synonym>
	<synonym relationship='synonym'>Raja bonaespeiensis Fowler, 1910</synonym>
	<synonym relationship='ambiguous synonym'>Raja capensis Müller &amp; Henle, 1841</synonym>
	<synonym relationship='misapplied name'>Raja gesneri (non Cuvier, 1829)</synonym>
	<synonym relationship='synonym'>Raja leiobatos Gronow, 1854</synonym>
	<synonym relationship='misapplied name'>Raja maderensis (non Lowe, 1838)</synonym>
	<synonym relationship='synonym'>Raja pontica Pallas, 1914</synonym>
	<synonym relationship='misapplied name'>Raja punctata (non Woodward, 1900-01)</synonym>
	<synonym relationship='synonym'>Raja rubus Bloch, 1784</synonym>
	<dcterms:created>1991-01-18</dcterms:created>
	<dcterms:modified>2017-09-09</dcterms:modified>
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	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=96325&amp;speccode=2059'>Miniconi, R.0 Complément à l'inventaire des poissons des eaux de Corse (1994-2011). Biologie - Pȇche - Localisation. Edition A Barcella Ajaccio, 50 p. (Ref. 96325)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=98752&amp;speccode=2059'>Sadio, O.0 Common names of some fish species in Senegal. Unpublished compilation of O. Sadio. (Ref. 98752)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=100834&amp;speccode=2059'>Barría, C., J. Navarro, M. Coll, U. Fernancez-Arcaya and R. Sáez-Liante0 Morphological parameters of abundant and threatened chondrichthyans of the northwestern Mediterranean Sea. J. Appl. Ichthyol. 31(1):114-119. (Ref. 100834)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=101110&amp;speccode=2059'>FAO-FIES0 Aquatic Sciences and Fisheries Information System (ASFIS) species list. Retrieved from http://www.fao.org/fishery/collection/asfis/en, [accessed 13/04/2015]. (Ref. 101110)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=102582&amp;speccode=2059'>Ulman, A., A Saad, K Zylich, D. Pauly and D. Zeller0 Reconstruction of Syria’s fisheries catches from 1950-2010: Signs of overexploitation. Acta Ichthyol. Piscat. 45(3):259-272. (Ref. 102582)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=105472&amp;speccode=2059'>Pihu, E.0 Loomade elu 4. köide. Kalad. Pihu, E. 1979. Loomade Elu, 4. Köide, Kalad. Valgus, Tallinn. (Ref. 105472)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=106276&amp;speccode=2059'>Wilhelms, I.0 Atlas of length-weight relationships of 93 fish and crustacean species from the North Sea and the North-East Atlantic (No. 12). Johann Heinrich von Thünen Institute, Federal Research Institute for Rural Areas, Forestry and Fisheries, 552 p. (Ref. 106276)</reference>
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	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=107742&amp;speccode=2059'>Barrientos-Medina, R.C., S. Avilés-Torres and J.A. Navarro-Alberto0 Length-weight relations of eight fish species from a small Caribbean coastal lagoon, Mexico. Acta Ichthyol. Piscat. 43(1):71-74. (Ref. 107742)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=107945&amp;speccode=2059'>Cheung, W.L., R. Watson and D. Pauly0 Signature of ocean warming in global fisheries catch. Nature 497:365-368. (Ref. 107945)</reference>
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	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=114953&amp;speccode=2059'>Last, P.R., W.T. White, M.R. de Carvalho, B. Séret, M.F.W. Stehmann and G.J.P. Naylor0 Rays of the world. CSIRO Publishing, Comstock Publishing Associates. i-ix + 1-790. (Ref. 114953)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=115039&amp;speccode=2059'>ICES0 Report of the Working Group on Elasmobranch Fishes (WGEF), 15-24 June 2016, Lisbon, Portugal. ICES CM/ACOM:20. 26 pp. (Ref. 115039)</reference>
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	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=115544&amp;speccode=2059'>Keskin, C.0 A preliminary study on demersal fishes in the southwestern Black Sea shelf (NW Turkey). J. Black Sea/Mediterranean Environment 18(3):341-349. (Ref. 115544)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=115876&amp;speccode=2059'>Crec'hriou, R., R. Neveu and P. Lenfant0 Length-weight relationship of main commercial fishes from the French Catalan coast. J. Appl. Ichthyol. 29:1191-1192. (Ref. 115876)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=116450&amp;speccode=2059'>Béarez, P., P. Pruvost, É. Feunteun, S. Iglésias, P. Francour, R. Causse, J. De Mazières, S. Tercerie and N. Bailly0 Checklist of the marine fishes from metropolitan France. Cybium 41(4):351-371. (Ref. 116450)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=117987&amp;speccode=2059'>Lynghammar, A., J.S. Christiansen, C.W. Mecklenburg, O.V. Karamushko, P.R. Møller and V.F. Gallucci0 Species richness and distribution of chondrichthyan fishes in the Arctic Ocean and adjacent seas. Biodiversity 14(1):57-66. (Ref. 117987)</reference>
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	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=118592&amp;speccode=2059'>Froese, R.0 Personal communication. (Table sent by the author). (Ref. 118592)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=118717&amp;speccode=2059'>Tsuboi, M., W. van der Bijl, B.T. Kopperud, J. Erritzoe, K.L. Voje, A. Kotrschal, K.E. Yopak, S.P. Collin, A.N. Iwaniuk and N. Kolm0 Breakdown of brain-body allometry and the encephalization of birds and mammals. Nat. Ecol. Evol. 2:1495-1500. (Ref. 118717)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=119290&amp;speccode=2059'>Kasapoglu, N. and E. Duzgunes0 Length-weight relationships of marine species caught by five gears from the Black Sea. Mediterranean Marine Science 15(1):95-100. (Ref. 119290)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=119547&amp;speccode=2059'>Mahé, K., E. Bellamy, J.-P. Delpech, C. Lazard, M. Salaun, Y. Vérin, F. Coppin and M. Travers-Trolet0 Evidence of a relationship between weight and total length of marine fish in the North-eastern Atlantic Ocean: physiological, spatial and temporal variations. Journal of the Marine Biological Association of the United Kingdom 98(3):627-625. (Ref. 119547)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=121381&amp;speccode=2059'>Demirhan, S.A., K. Seyhan and N. Basusta0 Dietary Overlap in Spiny Dogfish (&lt;i&gt;Squalus acanthias&lt;/i&gt;) and Thornback Ray (&lt;i&gt;Raja clavata&lt;/i&gt;) in the Southeastern Black Sea. Ekoloji 16(2007):1-8. (Ref. 121381)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=121645&amp;speccode=2059'>İşmen, A., C.Ç. Yiğın, M.A. İhsanoğlu and B. Daban0 Length-weight relationships for three elasmobranch species from the Sea of Marmara. J. Black Sea/Mediterranean Environment 24(1):65-73. (Ref. 121645)</reference>
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	<reference isbn='2350-4757' url='http://www.fishbase.org/references/FBRefSummary.php?id=128520&amp;speccode=2059'>Marčeta, B. and K. Pliberšek (eds)0 BiosWeb. Biological database of the Fisheries Research Institute of Slovenia. Ljubljana, Fisheries Research Institute of Slovenia, 2014. (www.biosweb.org. updated 13/04/2023). (Ref. 128520)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=129849&amp;speccode=2059'>dos Santos Cabral, S.0 &lt;i&gt;Raja&lt;/i&gt; spp. Spatial dynamics in the Arrábida Marine Park. Algrave University: Faculty of Science and Technology. Master of Science in Marine Biodiversity and Conservation, 60 p. (Ref. 129849)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=130435&amp;speccode=2059'>IUCN0 The IUCN Red List of Threatened Species. 1964 - onwards. https://www.iucnredlist.org/ (Ref. 130435)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=132125&amp;speccode=2059'>Ferragut-Perello, F., S. Ramirez-Amaro, A.C. Tsikliras, N. Petit-Marty, D. Dimarchopoulou0 Exploitation and conservation status of the thornback ray (&lt;i&gt;Raja clavata&lt;/i&gt;) in the Balearic Islands (Western Mediterranean). Fishes 8(2):117. (Ref. 132125)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=138364&amp;speccode=2059'>Zubillaga, C.P.0 High mislabeling and low levels of genetic diversity in skates from the Cantabrian Sea: a challenge for sustainable management in Rajidae. Spain: Oviedo, Universidad de Oviedo, MSc Thesis, 36 p. (Ref. 138364)</reference><dataObject>
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	<dc:description>50 cm wingspan; 3.2 kg; blackcolored snout Locality: Dingle, Kerry, Ireland.</dc:description>
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	<dc:description>Inhabits shelf and upper slope waters. Reported depth ranges varies, from 10-300 m (Ref. 4426,  82319, 88187), and from 300-577 m in the eastern Ionian Sea (Ref. 56504); most common in coastal waters between 10-60 m depth (Ref. 58137). Tolerates low salinities (Ref. 88171). Found on mud, sand and gravel bottoms, rarely on rougher bottoms (Ref. 6808, 58137). Nocturnal species. Feeds on all kinds of bottom animals, preferably crustaceans (Ref. 3167) and fishes (Ref. 114953). Undertakes migrations with mean distances of 54-117 km per month; shows a clear annual migration cycle (Ref. 89017, 89018), moves from deeper offshore waters (10-30 m) in autumn and winter to shallower areas (</dc:description>
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	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=27314&amp;speccode=2059'>McEachran, J.D. and K.A. Dunn 1998 Phylogenetic analysis of skates, a morphologically conservative clade of elasmobranchs (Chondrichthyes: Rajidae). Copeia 1998(2):271-290. (Ref. 27314)</reference></dataObject><dataObject>
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	<dc:description>Sub-rhomboid in shape with pointed wing-tips; disc-width 1,25 to 1,36 times in its length, its length 1,70 to 1,83 times in total length. Short rostrum, rounded at his extremity; pectoral fins with clear angles on lateral side; triangular pelvic fins (Ref. 39246). Dorsally prickly; large females also prickly throughout their ventral surface; juveniles and large males prickly along the margins of their discs and the underside of their snout. 30-50 thorns form a median row from the nape to the first dorsal fin; additional large 'buckler' thorns with swollen bases scattered on upper surface of disc in adults (Ref. 3167). Coloration is highly variable, with the dorsal surface having all shades of brown with dark and light spots and blotches; ventral surface white (Ref. 3167, 58137). </dc:description>
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	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=39246&amp;speccode=2059'>Capapé, C. and M. Desoutter 1981 Nouvelle description de &lt;i&gt;Raja (Raja) clavata&lt;/i&gt; Linnaeus, 1758 et note sur la validité de &lt;i&gt;Raja (Raja) capensis&lt;/i&gt; Müller et Henle, 1841. Cybium 5(4):23-39. (Ref. 39246)</reference></dataObject><dataObject>
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	<dc:description>Dorsal spines (total): 0;  Analspines: 0; Analsoft rays: 0</dc:description>
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	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=39246&amp;speccode=2059'>Capapé, C. and M. Desoutter 1981 Nouvelle description de &lt;i&gt;Raja (Raja) clavata&lt;/i&gt; Linnaeus, 1758 et note sur la validité de &lt;i&gt;Raja (Raja) capensis&lt;/i&gt; Müller et Henle, 1841. Cybium 5(4):23-39. (Ref. 39246)</reference></dataObject><dataObject>
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	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=114953&amp;speccode=2059'>Last, P.R., W.T. White, M.R. de Carvalho, B. Séret, M.F.W. Stehmann and G.J.P. Naylor 2016 Rays of the world. CSIRO Publishing, Comstock Publishing Associates. i-ix + 1-790. (Ref. 114953)</reference></dataObject><dataObject>
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	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=4931&amp;speccode=2059'>FAO 1992 FAO yearbook. Fishery statistics: catches and landings. Vol. 70: 1990. FAO Fish. Ser. 38:647 p. (Ref. 4931)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=4699&amp;speccode=2059'>International Game Fish Association 1991 World record game fishes. International Game Fish Association, Florida, USA. (Ref. 4699)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=274&amp;speccode=2059'>Flower, S.S. 1935 Further notes on the duration of life in animals. I. Fishes: as determined by otolith and scale-readings and direct observations on living individuals. Proc. Zool. Soc. London 2:265-304. (Ref. 274)</reference></dataObject><dataObject>
	<dc:identifier>FB-LifeCycle-2059</dc:identifier>
	<dataType>http://purl.org/dc/dcmitype/Text</dataType>
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	<agent homepage='' logoURL='' role='recorder'>Cristina V. Garilao</agent>
	<dcterms:created>1991-01-18 00:00:00</dcterms:created>
	<dcterms:modified>2017-09-09 00:00:00</dcterms:modified>
	<dc:title></dc:title>
	<dc:language></dc:language>
	<license>http://creativecommons.org/licenses/by-nc/3.0/</license>
	<dc:rights></dc:rights>
	<dcterms:bibliographicCitation></dcterms:bibliographicCitation>
	<dc:source>http://www.fishbase.org//Reproduction/FishReproSummary.php?ID=2059&amp;StockCode=2254</dc:source>
	<dc:description>Oviparous. Distinct pairing with embrace. Polyandrous species (Ref. 89024). Paired eggs are laid and deposited on shallow sand, mud, pebble or gravel bottoms (Ref. 205, 89025). Up to 170 egg cases can be laid by a single female in a year (Ref. 3167), but average fecundity is much lower (around 48-74 eggs) (Ref. 3603, 31302). In northwestern Europe, egg cases are laid during spring (Ref. 3167) and in the Mediterranean during winter and spring (Ref. 3167). Egg cases are oblong capsules with stiff pointed horns at the corners, each containing one embryo (Ref. 205). Capsules are 5.0-9.0 cm long without the horns (Ref. 41250, 88187) and 3.4-6.8 cm wide (Ref. 41250). Egg cases are anchored with an adhesive film (Ref. 82399). Embryos feed solely on yolk (Ref. 50449). Egg cases hatch after about 4-5 months and pups are about 11-13 cm TL (Ref. 88864). Mating season from February to September, peaking in June (Ref. 74501). Adults observed to form same-sex aggregations during the mating season with females moving to shallower inshore waters approximately a month before the males (Ref 3603, 58137, 74501). Mating does not occur in the Baltic Sea (Ref. 82311).</dc:description>
	<mediaURL></mediaURL>
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	<location></location>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=205&amp;speccode=2059'>Breder, C.M. and D.E. Rosen 1966 Modes of reproduction in fishes. T.F.H. Publications, Neptune City, New Jersey. 941 p. (Ref. 205)</reference></dataObject><dataObject>
	<dc:identifier>FB-TrophicStrategy-2059</dc:identifier>
	<dataType>http://purl.org/dc/dcmitype/Text</dataType>
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	<agent homepage='' logoURL='' role='recorder'>Pascualita Sa-a</agent>
	<dcterms:created>1992-12-14 00:00:00</dcterms:created>
	<dcterms:modified>2012-01-12 20:46:55</dcterms:modified>
	<dc:title></dc:title>
	<dc:language></dc:language>
	<license>http://creativecommons.org/licenses/by-nc/3.0/</license>
	<dc:rights></dc:rights>
	<dcterms:rightsHolder>WorldFish Center - FishBase</dcterms:rightsHolder>
	<dcterms:bibliographicCitation></dcterms:bibliographicCitation>
	<audience>General public</audience>
	<dc:source>clavata</dc:source>
	<subject>http://rs.tdwg.org/ontology/voc/SPMInfoItems#Threats</subject>
	<dc:description>Nocturnal predator. Feeds on all kinds of bottom animals (Ref. 3167). Polychaetes and crustaceans are important prey items of smaller individuals, bony fishes become more important prey items of larger individuals (Ref. 28070). </dc:description>
	<mediaURL></mediaURL>
	<thumbnailURL></thumbnailURL>
	<location></location>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=4921&amp;speccode=2059'>Ajayi, T.O. 1982 Food and feeding habits of &lt;i&gt;Raja&lt;/i&gt; species (Batoidei) in Carmarthen Bay, Bristol Channel. J. Mar. Biol. Ass. U.K. 62:215-223. (Ref. 4921)</reference></dataObject><dataObject>
	<dc:identifier>FB-Size-2059</dc:identifier>
	<dataType>http://purl.org/dc/dcmitype/Text</dataType>
	<mimeType>text/html</mimeType>
	<agent homepage='' logoURL='' role='recorder'>Susan M. Luna</agent>
	<dcterms:created>1991-01-18 00:00:00</dcterms:created>
	<dcterms:modified>2017-09-09 00:00:00</dcterms:modified>
	<dc:title>Max. size</dc:title>
	<dc:language></dc:language>
	<license>http://creativecommons.org/licenses/by-nc/3.0/</license>
	<dc:rights></dc:rights>
	<dcterms:rightsHolder>WorldFish Center - FishBase</dcterms:rightsHolder>
	<dcterms:bibliographicCitation></dcterms:bibliographicCitation>
	<audience>General public</audience>
	<dc:source>http://www.fishbase.org/summary/SpeciesSummary.php?id=2059</dc:source>
	<subject>http://rs.tdwg.org/ontology/voc/SPMInfoItems#Habitat</subject>
	<dc:description>105 cm TL (male/unsexed; (Ref. 53748)); 139 cm TL (female); max. published weight: 18.0 kg (Ref. 35388); max. reported age: 15 years (Ref. 41305)</dc:description>
	<mediaURL></mediaURL>
	<thumbnailURL></thumbnailURL>
	<location></location>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=53748&amp;speccode=2059'>Sulikowski, J.A., J. Kneebone, S. Elzey, J. Jurek, P.D. Danley, W. Huntting Howell and P.C.W. Tsang 2005 Age and growth estimates of the thorny skate (&lt;i&gt;Amblyraja radiata&lt;/i&gt;) in the western Gulf of Maine. Fish. Bull. 103:161-168. (Ref. 53748)</reference>
	<reference isbn='87-90787-00-5' url='http://www.fishbase.org/references/FBRefSummary.php?id=35388&amp;speccode=2059'>Muus, B.J. and J.G. Nielsen 1999 Sea fish. Scandinavian Fishing Year Book, Hedehusene, Denmark. 340 p. (Ref. 35388)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=41305&amp;speccode=2059'>Walker, P.A. 1998 Fleeting images. Dynamics of North Sea ray populations. University of Amsterdam. 145 p. Ph.D. dissertation. (Ref. 41305)</reference></dataObject><dataObject>
	<dc:identifier>FB-Threats-2</dc:identifier>
	<dataType>http://purl.org/dc/dcmitype/Text</dataType>
	<mimeType>text/html</mimeType>
	<agent homepage='' logoURL='' role='recorder'>Belen Acosta</agent>
	<dcterms:created>1990-10-18 00:00:00</dcterms:created>
	<dcterms:modified>2024-03-07 00:00:00</dcterms:modified>
	<dc:title></dc:title>
	<dc:language></dc:language>
	<license>http://creativecommons.org/licenses/by-nc/3.0/</license>
	<dc:rights></dc:rights>
	<dcterms:rightsHolder>WorldFish Center - FishBase</dcterms:rightsHolder>
	<dcterms:bibliographicCitation></dcterms:bibliographicCitation>
	<audience>General public</audience>
	<dc:source>http://www.fishbase.org/summary/SpeciesSummary.php?id=2</dc:source>
	<subject>http://rs.tdwg.org/ontology/voc/SPMInfoItems#Distribution</subject>
	<dc:description>Least Concern (LC) , IUCN Grouper and Wrasse Specialist Group</dc:description>
	<mediaURL></mediaURL>
	<thumbnailURL></thumbnailURL>
	<location></location>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=57073'>IUCN 2006 2006 IUCN red list of threatened species. www.iucnredlist.org. Downloaded July 2006.</reference></dataObject><dataObject>
	<dc:identifier>FB-Habitat-2059</dc:identifier>
	<dataType>http://purl.org/dc/dcmitype/Text</dataType>
	<mimeType>text/html</mimeType>
	<agent homepage='' logoURL='' role='recorder'>Susan M. Luna</agent>
	<dcterms:created>1991-01-18 00:00:00</dcterms:created>
	<dcterms:modified>2017-09-09 00:00:00</dcterms:modified>
	<dc:title>Environment</dc:title>
	<dc:language></dc:language>
	<license>http://creativecommons.org/licenses/by-nc/3.0/</license>
	<dc:rights></dc:rights>
	<dcterms:bibliographicCitation></dcterms:bibliographicCitation>
	<dc:source>http://www.fishbase.org/summary/SpeciesSummary.php?id=2059</dc:source>
	<dc:description>demersal; marine; depth range 5 - 1020 m (Ref. 114953)</dc:description>
	<mediaURL></mediaURL>
	<thumbnailURL></thumbnailURL>
	<location></location>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=114953&amp;speccode=2059'>Last, P.R., W.T. White, M.R. de Carvalho, B. Séret, M.F.W. Stehmann and G.J.P. Naylor 2016 Rays of the world. CSIRO Publishing, Comstock Publishing Associates. i-ix + 1-790. (Ref. 114953)</reference></dataObject>
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