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	<dc:identifier>FB-15868</dc:identifier>
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	<dwc:Kingdom>Animalia</dwc:Kingdom>
	<dwc:Phylum>Chordata</dwc:Phylum>
	<dwc:Class>Teleostei</dwc:Class>
	<dwc:Order>Gobiiformes</dwc:Order>
	<dwc:Family>Gobiidae</dwc:Family>
	<dwc:Genus>Gymnogobius</dwc:Genus>
	<dwc:ScientificName>Gymnogobius heptacanthus (Hilgendorf, 1879)</dwc:ScientificName>
	<commonName xml:lang='ja'>Isaza</commonName>
	<commonName xml:lang='ja'>イサザ</commonName>
	<commonName xml:lang=''>疣舌裸头鰕虎鱼</commonName>
	<commonName xml:lang=''>疣舌裸頭鰕虎魚</commonName>
	<synonym relationship='synonym'>Chaenogobius isaza Tanaka, 1916</synonym>
	<dcterms:created>1995-12-04</dcterms:created>
	<dcterms:modified>2024-05-22</dcterms:modified>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=559&amp;speccode=15869'>Masuda, H., K. Amaoka, C. Araga, T. Uyeno and T. Yoshino0 The fishes of the Japanese Archipelago. Vol. 1. Tokai University Press, Tokyo, Japan. 437 p. (text). (Ref. 559)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=4517&amp;speccode=15869'>Hureau, J.-C.0 La base de données GICIM: Gestion informatisée des collections ichthyologiques du Muséum. p. 225-227. In Atlas Préliminaire des Poissons d'Eau Douce de France. Conseil Supérieur de la Pêche, Ministère de l'Environnement, CEMAGREF et Muséum national d'Histoire naturelle, Paris. (Ref. 4517)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=8973&amp;speccode=15869'>Vasil'ev, V.P.0 Chromosome numbers in fish-like vertebrates and fish. J. Ichthyol. 20(3):1-38. (Ref. 8973)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=9902&amp;speccode=15869'>Okiyama, M.0 An atlas of the early stage fishes in Japan. Tokai University Press, Tokyo, Japan. 1154 p. (Ref. 9902)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=11344&amp;speccode=15869'>Bauchot, M.-L., J.-M. Ridet and M. Diagne0 Encephalization in Gobioidei (Teleostei). Jap. J. Ichthyol. 36(1):63-74. (Ref. 11344)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=30163&amp;speccode=15869'>Arai, R. and Y. Sawada0 Chromosomes of Japanese gobioid fishes (III). Bull. Natn. Sci. Mus., Ser.A. 1(4):225-232. Tokyo. (Ref. 30163)</reference>
	<reference isbn='957-8596-59-6' url='http://www.fishbase.org/references/FBRefSummary.php?id=31517&amp;speccode=15869'>Wu, H.L., K.-T. Shao and C.F. Lai (eds.)0 Latin-Chinese dictionary of fishes names. The Sueichan Press, Taiwan. 1028 p. (Ref. 31517)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=34370&amp;speccode=15869'>Klinkhardt, M, M. Tesche and H. Greven0 Database of fish chromosomes. Westarp Wissenschaften. (Ref. 34370)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=41827&amp;speccode=15869'>Anonymous0 Freshwater fish collection database of the National Science Museum. National Science Museum, 3-23-1 Hyakunin-cho, Shinjuku-ku, Tokyo 169-0073, Japan. (Ref. 41827)</reference>
	<reference isbn='4-486-01570-3' url='http://www.fishbase.org/references/FBRefSummary.php?id=43239&amp;speccode=15869'>Nakabo, T.0 Fishes of Japan with pictorial keys to the species, English edition II. Tokai University Press, Japan, pp 867-1749. (Ref. 43239)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=52995&amp;speccode=15869'>Morey, G., J. Moranta, E. Massutí, A. Grau, M. Linde, F. Riera and B. Morales-Nin0 Weight-length relationships of littoral to lower slope fishes from the western Mediterranean. Fish. Res. 62(1):89-96. (Ref. 52995)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=58108&amp;speccode=15869'>Chinese Academy of Fishery Sciences0 Chinese aquatic germplasm resources database. http://zzzy.fishinfo.cn (Ref. 58108)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=76911&amp;speccode=15869'>Nakazawa, T., N. Ishida, M. Kato and N. Yamamura0 Larger body size with higher predation rate. Ecol. Freshwat. Fish16(3):362-372. (Ref. 76911)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=76913&amp;speccode=15869'>Takahashi, D., H. Asada, T. Takeyama, M. Takahata, R. Katoh, S. Awata and M. Kohda0 Why egg-caring males of Isaza (&lt;i&gt;Gymnogobius isaza&lt;/i&gt;, Gobiidae) refuse additional females: preliminary field observations. J. Ethol. 22(2):153-159. (Ref. 76913)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=76915&amp;speccode=15869'>Toshitaka, H. and S. Takahashi0 Reproductive strategy and interspecific competition in the lake-living gobiid fish isaza, &lt;i&gt;Chaenogobius isaza&lt;/i&gt;. J. Ethol. 5(2):185-196. (Ref. 76915)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=78994&amp;speccode=15869'>Fisheries Management Division of the Shiga Prefecture Agriculture and Fisheries Department0 Shiga's Fisheries (Fiscal Year 2007). Shiga no Suisan (Heisei 19 Nendo), pp. 77-79. (Ref. 78994)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=79825&amp;speccode=15869'>Wildlife Survey Group in Shiga Prefecture (ed.)0 Red data book of Shiga Prefecture. Sunrise Press, Shiga, p. 475. (Ref. 79825)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=79886&amp;speccode=15869'>Uonokai, T. Nakajima and K. Ohara (eds.)0 Activities of Uonokai (Fish survey group): Fish distribution in the watershed of Lake Biwa. Shiga, Japan: Lake Biwa Museum. 233 p. (Ref. 79886)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=81467&amp;speccode=15869'>Stevenson, D.E.0 Systematics and distribution of fishes of the Asian goby genera &lt;i&gt;Chaenogobius&lt;/i&gt; and &lt;i&gt;Gymnogobius&lt;/i&gt; (Osteichthyes: Perciformes: Gobiidae), with the description of a new species. Spec. Divers. 7(3):251-312. (Ref. 81467)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=95975&amp;speccode=15869'>Shiga Prefecture0 Red Data Book of Shiga Prefecture 2010. Shiga Prefecture, Otsu. (http://www.pref.shiga.lg.jp/d/shizenkankyo/kyoseijourei.html; http://www.pref.shiga.lg.jp/d/shizenkankyo/files/risuto.xls) (Ref. 95975)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=118717&amp;speccode=15869'>Tsuboi, M., W. van der Bijl, B.T. Kopperud, J. Erritzoe, K.L. Voje, A. Kotrschal, K.E. Yopak, S.P. Collin, A.N. Iwaniuk and N. Kolm0 Breakdown of brain-body allometry and the encephalization of birds and mammals. Nat. Ecol. Evol. 2:1495-1500. (Ref. 118717)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=122096&amp;speccode=15869'>Nakazawa, T., C.-H. Tsai, N. Ishida and M. Kato0 Annual variations in length-weight relationship of &lt;i&gt;Gymnogobius isaza&lt;/i&gt; (Tanaka, 1916) from Lake Biwa, Japan. J. Appl. Ichthyol. 27:1135-1136. (Ref. 122096)</reference><dataObject>
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	<agent homepage='http://www.fishbase.de/rfroese' logoURL='' role='recorder'>Rainer Froese</agent>
	<dcterms:created>1995-12-04 00:00:00</dcterms:created>
	<dcterms:modified>2021-07-22 00:00:00</dcterms:modified>
	<dc:title>Biology</dc:title>
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	<dc:description>Inhabit offshore areas in the lake almost throughout the year (Ref. 76915).  Spawn in spring (Ref. 76915).  Semelparous (Ref. 76911).  Males care for the eggs in nest (Ref. 76913).  Caught for food (Ref. 559).</dc:description>
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	<reference isbn='4-486-01570-3' url='http://www.fishbase.org/references/FBRefSummary.php?id=43239&amp;speccode=15869'>Nakabo, T. 2002 Fishes of Japan with pictorial keys to the species, English edition II. Tokai University Press, Japan, pp 867-1749. (Ref. 43239)</reference></dataObject><dataObject>
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	<agent homepage='' logoURL='' role='recorder'>Armi G. Torres</agent>
	<dcterms:created>2008-07-02 00:00:00</dcterms:created>
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	<dc:description>Dorsal spines (total): 7; Dorsal soft rays (total): 10;  Analspines: 1; Analsoft rays: 10</dc:description>
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	<reference isbn='4-486-01570-3' url='http://www.fishbase.org/references/FBRefSummary.php?id=43239&amp;speccode=15869'>Nakabo, T. 2002 Fishes of Japan with pictorial keys to the species, English edition II. Tokai University Press, Japan, pp 867-1749. (Ref. 43239)</reference></dataObject><dataObject>
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	<dc:description>Asia:  endemic to Lake Biwa, Japan.  Introduced to Lakes Kasumigaura and Sagami.</dc:description>
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	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=559&amp;speccode=15869'>Masuda, H., K. Amaoka, C. Araga, T. Uyeno and T. Yoshino 1984 The fishes of the Japanese Archipelago. Vol. 1. Tokai University Press, Tokyo, Japan. 437 p. (text). (Ref. 559)</reference></dataObject><dataObject>
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	<dc:title>Importance</dc:title>
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	<dc:description>fisheries: minor commercial</dc:description>
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	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=559&amp;speccode=15869'>Masuda, H., K. Amaoka, C. Araga, T. Uyeno and T. Yoshino 1984 The fishes of the Japanese Archipelago. Vol. 1. Tokai University Press, Tokyo, Japan. 437 p. (text). (Ref. 559)</reference></dataObject><dataObject>
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	<dc:description>Migrate to the shore to spawn under stones in spring (Ref. 76915).  Males have only two brood cycles, mating with the same female during each breeding cycle (Ref. 76915).</dc:description>
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	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=76915&amp;speccode=15869'>Toshitaka, H. and S. Takahashi 1987 Reproductive strategy and interspecific competition in the lake-living gobiid fish isaza, &lt;i&gt;Chaenogobius isaza&lt;/i&gt;. J. Ethol. 5(2):185-196. (Ref. 76915)</reference></dataObject><dataObject>
	<dc:identifier>FB-Size-15869</dc:identifier>
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	<dc:title>Max. size</dc:title>
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	<dc:description>7.0 cm SL (male/unsexed; (Ref. 43239))</dc:description>
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	<reference isbn='4-486-01570-3' url='http://www.fishbase.org/references/FBRefSummary.php?id=43239&amp;speccode=15869'>Nakabo, T. 2002 Fishes of Japan with pictorial keys to the species, English edition II. Tokai University Press, Japan, pp 867-1749. (Ref. 43239)</reference></dataObject><dataObject>
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	<dc:description>Least Concern (LC) , IUCN Grouper and Wrasse Specialist Group</dc:description>
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	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=57073'>IUCN 2006 2006 IUCN red list of threatened species. www.iucnredlist.org. Downloaded July 2006.</reference></dataObject><dataObject>
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	<dc:description>pelagic; non-migratory; freshwater</dc:description>
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