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<taxon>
	<dc:identifier>FB-1118</dc:identifier>
	<dc:source>http://www.fishbase.org/summary/SpeciesSummary.php?id=1118</dc:source>
	<dwc:Kingdom>Animalia</dwc:Kingdom>
	<dwc:Phylum>Chordata</dwc:Phylum>
	<dwc:Class>Teleostei</dwc:Class>
	<dwc:Order>Acanthuriformes</dwc:Order>
	<dwc:Family>Pomacanthidae</dwc:Family>
	<dwc:Genus>Pomacanthus</dwc:Genus>
	<dwc:ScientificName>Pomacanthus paru (Bloch, 1787)</dwc:ScientificName>
	<commonName xml:lang='pl'>Ustniczek francuzik</commonName>
	<commonName xml:lang='pt'>Enxada</commonName>
	<commonName xml:lang='pt'>Frade</commonName>
	<commonName xml:lang='pt'>Gordinho</commonName>
	<commonName xml:lang='pt'>Jandaia</commonName>
	<commonName xml:lang='pt'>Mercador</commonName>
	<commonName xml:lang='pt'>Paru-preto</commonName>
	<commonName xml:lang='pt'>Paru</commonName>
	<commonName xml:lang='pt'>Paru-beija-moça</commonName>
	<commonName xml:lang='pt'>Paru-bordado</commonName>
	<commonName xml:lang='pt'>Paru-da-pedra</commonName>
	<commonName xml:lang='pt'>Parú-da-pedra</commonName>
	<commonName xml:lang='pt'>Paru-dourado</commonName>
	<commonName xml:lang='pt'>Paru-frade</commonName>
	<commonName xml:lang='pt'>Paru-listrado</commonName>
	<commonName xml:lang='pt'>Parumbeba</commonName>
	<commonName xml:lang='pt'>Parum-dourado</commonName>
	<commonName xml:lang='pt'>Peixe-frade</commonName>
	<commonName xml:lang='pt'>Perambeba</commonName>
	<commonName xml:lang='pt'>Sambuio</commonName>
	<commonName xml:lang='da'>Grå kejserfisk</commonName>
	<commonName xml:lang='de'>Großflossen-Kaiserfisch</commonName>
	<commonName xml:lang='en'>Angelfish</commonName>
	<commonName xml:lang='en'>Gray angelfish</commonName>
	<commonName xml:lang='en'>Gray Angelfish</commonName>
	<commonName xml:lang='en'>Grey angelfish</commonName>
	<commonName xml:lang='en'>Pot cover</commonName>
	<commonName xml:lang='et'>Hall ingelkala</commonName>
	<commonName xml:lang='fr'>Demoiselle blanche</commonName>
	<commonName xml:lang='ja'>Nezumi-yakko</commonName>
	<commonName xml:lang='ru'>рыба-ангел серая</commonName>
	<commonName xml:lang='sv'>Svart Kejsarfisk</commonName>
	<commonName xml:lang='es'>Cachama blanca</commonName>
	<commonName xml:lang='es'>Chivirica gris</commonName>
	<commonName xml:lang='es'>Gallineta café</commonName>
	<commonName xml:lang='es'>Guinea</commonName>
	<commonName xml:lang='es'>Isabelita</commonName>
	<commonName xml:lang='es'>Isabelita blanca</commonName>
	<commonName xml:lang='es'>Manai oscuro</commonName>
	<commonName xml:lang=''>Francoski cesarček</commonName>
	<commonName xml:lang=''>Palanache</commonName>
	<synonym relationship='misapplied name'>Apolemichthys arcuatus (non Gray, 1831)</synonym>
	<synonym relationship='synonym'>Chaetodon arcuatus Linnaeus, 1758</synonym>
	<synonym relationship='synonym'>Chaetodon littoricola Poey, 1868</synonym>
	<synonym relationship='synonym'>Chaetodon lutescens Bonnaterre, 1788</synonym>
	<synonym relationship='synonym'>Chaetodon quinquecinctus Cuvier, 1829</synonym>
	<synonym relationship='synonym'>Chetodon lutescens Bonnaterre, 1788</synonym>
	<synonym relationship='misapplied name'>Holacanthus arcuatus (non Gray, 1831)</synonym>
	<synonym relationship='synonym'>Pomacanthus baltcatus Cuvier, 1831</synonym>
	<synonym relationship='synonym'>Pomacanthus balteatus Cuvier, 1831</synonym>
	<synonym relationship='synonym'>Pomacanthus cingulatus Cuvier, 1831</synonym>
	<synonym relationship='synonym'>Pomacanthus cinquecinctus Cuvier, 1829</synonym>
	<dcterms:created>1990-10-17</dcterms:created>
	<dcterms:modified>2022-10-20</dcterms:modified>
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	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=57616&amp;speccode=1117'>Dominici-Arosemena, A. and M. Wolff0 Reef fish community structure in Bocas del Toro (Caribbean, Panamá): Gradients in habitat complexity and exposure. Caribbean J. Sci. 41(3):613-637. (Ref. 57616)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=57756&amp;speccode=1117'>Floeter, S.R., J.L. Gasparini, L.A. Rocha, C.E.L. Ferreira, C.A. Rangel and B.M. Feitoza0 Brazilian reef fish fauna: checklist and remarks (updated Jan. 2003). Brazilian Reef Fish Project: www.brazilianreeffish.cjb.net. (Ref. 57756)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=74908&amp;speccode=1117'>Vega-Cendejas, M.E.0 Ictiofauna de la reserva de la biosfera Celestún, Yucatán: una contribución al conocimiento de su biodiversidad. Universidad Nacional Autónoma de México. Anales del Instituto de Biología, Serie Zoología 75:193-206. (Ref. 74908)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=78464&amp;speccode=1117'>McEachran, J.D. and J.D. Fechhelm0 Fishes of the Gulf of Mexico. Vol. 2. Austin : University of Texas Press. viii+1004p. (Ref. 78464)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=80860&amp;speccode=1117'>Menezes, N.A., P.A. Buckup, J.L. Figueiredo and R.L. Moura0 Catálogo das espécies de peixes marinhos do Brasil. São Paulo. Museu de Zoologia da Universidade de São Paulo, 160 pp. (Ref. 80860)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=81979&amp;speccode=1117'>Sampaio, C.L.S. and M.C. Nottingham0 Guia para identificação de peixes ornamentais - volume I: espécies marinhas. Brasília: Ibama, 205 p. (Ref. 81979)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=84211&amp;speccode=1117'>Feitoza, B.M., R.S. Rosa and L.A. Rocha0 Ecology and zoogeography of deep-reef fishes in Northeastern Brazil. Bull. Mar. Sci. 76(3):725-742. (Ref. 84211)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=86870&amp;speccode=1117'>Eschmeyer, W.N. and Fricke R. (eds.)0 Catalog of fishes. Updated internet version of 05 May 2011. Catalog databases of CAS cited in FishBase (website). (Ref. 86870)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=87232&amp;speccode=1117'>Belize Tropical Research and Education Center0 Species lists. https://www.belizemarinetrec.com/links/ [Accessed 27/03/2025]. (Ref. 87232)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=101194&amp;speccode=1117'>Patzner, R.A.0 Reproductive strategies of fish. pp. 311-350.  In  Rocha, M.J., A. Arukwe and B.G. Kapoor (eds). Fish reproduction: cytology, biology and ecology. Science Publisher, Inc. Oxford. 631 p. (Ref. 101194)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=105472&amp;speccode=1117'>Pihu, E.0 Loomade elu 4. köide. Kalad. Pihu, E. 1979. Loomade Elu, 4. Köide, Kalad. Valgus, Tallinn. (Ref. 105472)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=126840&amp;speccode=1117'>Simon, T., H.T. Pinheiro, R.L. Moura, A. Carvalho-Filho, L.A. Rocha, A.S. Martins, E. Mazzei, R.B. Francini-Filho, G.M. Amado-Filho and J.-C. Joyeux0 Mesophotic fishes of the Abrolhos Shelf, the largest reef ecosystem in the South Atlantic. J. Fish Biol. 89:990-1001. (Ref. 126840)</reference>
	<reference isbn='2350-4757' url='http://www.fishbase.org/references/FBRefSummary.php?id=128520&amp;speccode=1117'>Marčeta, B. and K. Pliberšek (eds)0 BiosWeb. Biological database of the Fisheries Research Institute of Slovenia. Ljubljana, Fisheries Research Institute of Slovenia, 2014. (www.biosweb.org. updated 13/04/2023). (Ref. 128520)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=128811&amp;speccode=1117'>Harborne, A.0 Fish and coral species lists compiled by Coral Cay Conservation:  Belize 1990-1998. Coral Cay Conservation, 26p. (Ref. 128811)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=137577&amp;speccode=1117'>Price, S.A., S.T. Friedman, K.A. Corn, O. Larouche, K. Brockelsby, A.J. Lee, M. Nagaraj, N.G. Bertrand, M. Danao, M.C. Coyne, J.R. Estrada, R. Friedman, E. Hoeft, M. Iwan, D. Gross, J.H. Kao, B. Landry, M.J. Linares, C. McGlinn and …0 FishShapes v1: Functionally relevant measurements of teleost shape and size on three dimensions. Ecology 103(e3829):1-2. (Ref. 137577)</reference><dataObject>
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	<dc:description>Maximum depth from Ref. 126840.  Common in coral reefs, usually solitary, occasionally in pairs.  Juveniles are part-time cleaners.  Feed mainly on sponges, but also takes tunicates, algae, zoantharians, gorgonians, hydroids, bryozoans, and seagrasses.  Oviparous (Ref. 240).  Monogamous and always in home-ranging pairs, that is they do not defend an exclusive territory (Ref. 52884).  Flesh reported to be of excellent quality (Ref. 3797); marketed fresh and salted (Ref. 5217).  Friendly toward divers (Ref. 9710).  Have been reared in captivity (Ref. 35419).</dc:description>
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	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=4858&amp;speccode=1117'>Allen, G.R. 1985 Butterfly and angelfishes of the world. Vol. 2. 3rd edit. in English. Mergus Publishers, Melle, Germany. (Ref. 4858)</reference></dataObject><dataObject>
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	<dc:description>Pale gray around mouth, and pale gray margin on caudal fin.  Inside of pectoral fin yellow (Ref. 26938).  Juveniles are black with two light yellow bars on body and three on head; caudal fin yellow with a vertically elongate, nearly rectangular or hemispherical black spot in middle (Ref. 13442).</dc:description>
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	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=4858&amp;speccode=1117'>Allen, G.R. 1985 Butterfly and angelfishes of the world. Vol. 2. 3rd edit. in English. Mergus Publishers, Melle, Germany. (Ref. 4858)</reference></dataObject><dataObject>
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	<dcterms:created>1994-06-21 00:00:00</dcterms:created>
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	<dc:description>Dorsal spines (total): 9; Dorsal soft rays (total): 31 - 33;  Analspines: 3; Analsoft rays: 23 - 25</dc:description>
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	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=4858&amp;speccode=1117'>Allen, G.R. 1985 Butterfly and angelfishes of the world. Vol. 2. 3rd edit. in English. Mergus Publishers, Melle, Germany. (Ref. 4858)</reference></dataObject><dataObject>
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	<dc:description>Western Atlantic:  New England, USA to the vicinity of Rio de Janeiro, Brazil, including the Gulf of Mexico and the Caribbean (Ref. 26938).</dc:description>
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	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=4858&amp;speccode=1117'>Allen, G.R. 1985 Butterfly and angelfishes of the world. Vol. 2. 3rd edit. in English. Mergus Publishers, Melle, Germany. (Ref. 4858)</reference></dataObject><dataObject>
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	<dc:title>Importance</dc:title>
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	<dc:description>fisheries: minor commercial; aquarium: commercial; price category: high; price reliability: very questionable: based on ex-vessel price for species in this family</dc:description>
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	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=6077&amp;speccode=1117'>Cervigón, F. and W. Fischer 1979 INFOPESCA. Catálogo de especies marinas de interes economico actual o potencial para América Latina. Parte 1. Atlántico centro y suroccidental. FAO/UNDP, SIC/79/1. 372 p. FAO, Rome. (Ref. 6077)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=4858&amp;speccode=1117'>Allen, G.R. 1985 Butterfly and angelfishes of the world. Vol. 2. 3rd edit. in English. Mergus Publishers, Melle, Germany. (Ref. 4858)</reference></dataObject><dataObject>
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	<dc:description>Monogamous mating is observed as both facultative and social (Ref. 52884).</dc:description>
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	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=240&amp;speccode=1117'>Thresher, R.E. 1984 Reproduction in reef fishes. T.F.H. Publications, Inc. Ltd., Neptune City, New Jersey. 399 p. (Ref. 240)</reference></dataObject><dataObject>
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	<agent homepage='' logoURL='' role='recorder'>Pascualita Sa-a</agent>
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	<dc:source>arcuatus</dc:source>
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	<dc:description>Common in coral reefs.  Feed mainly on sponges, but also takes tunicates, algae, zoantharians, gorgonians, hydroids, bryozoans, and seagrasses.  Omnivore (Ref. 57616).  Sessile invertebrate feeder (Ref. 126840).</dc:description>
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	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=33'>Randall, J.E. 1967 Food habits of reef fishes of the West Indies. Stud. Trop. Oceanogr. Miami 5:665-847.</reference></dataObject><dataObject>
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	<dc:title>Max. size</dc:title>
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	<dc:source>http://www.fishbase.org/summary/SpeciesSummary.php?id=1117</dc:source>
	<dc:description>60.0 cm TL (male/unsexed; (Ref. 3797)); max. published weight: 1,830 g (Ref. 40637)</dc:description>
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	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=3797&amp;speccode=1117'>Burgess, W.E. 1978 Pomacanthidae. In W. Fischer (ed.) FAO species identification sheets for fishery purposes. Western Central Atlantic (Fishing Area 31). Vol. 3. [var. pag.] FAO, Rome. (Ref. 3797)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=40637&amp;speccode=1117'>IGFA 2001 Database of IGFA angling records until 2001. IGFA, Fort Lauderdale, USA. (Ref. 40637)</reference></dataObject><dataObject>
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	<agent homepage='' logoURL='' role='recorder'>Belen Acosta</agent>
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	<dc:description>Least Concern (LC) , IUCN Grouper and Wrasse Specialist Group</dc:description>
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	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=57073'>IUCN 2006 2006 IUCN red list of threatened species. www.iucnredlist.org. Downloaded July 2006.</reference></dataObject><dataObject>
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	<agent homepage='' logoURL='' role='recorder'>Susan M. Luna</agent>
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	<dc:description>reef-associated; non-migratory (Ref. 52884); marine; depth range 2 - 60 m (Ref. 9626)</dc:description>
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	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=52884&amp;speccode=1117'>Whiteman, E.A. and I.M. Côté 2004 Monogamy in marine fishes. Biol. Rev. 79:351-375. (Ref. 52884)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=9626&amp;speccode=1117'>Cervigón, F. 1993 Los peces marinos de Venezuela. Volume 2. Fundación Científica Los Roques, Caracas,Venezuela. 497 p. (Ref. 9626)</reference></dataObject>
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