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	<dc:identifier>FB-10675</dc:identifier>
	<dc:source>http://www.fishbase.org/summary/SpeciesSummary.php?id=10675</dc:source>
	<dwc:Kingdom>Animalia</dwc:Kingdom>
	<dwc:Phylum>Chordata</dwc:Phylum>
	<dwc:Class>Teleostei</dwc:Class>
	<dwc:Order>Cichliformes</dwc:Order>
	<dwc:Family>Cichlidae</dwc:Family>
	<dwc:Genus>Coptodon</dwc:Genus>
	<dwc:ScientificName>Coptodon gutturosus (Stiassny, Schliewen &amp; Dominey, 1992)</dwc:ScientificName>
	<commonName xml:lang=''>大咽非鯽</commonName>
	<commonName xml:lang=''>大咽非鲫</commonName>
	<synonym relationship='synonym'>Coptodon gutturosa (Stiassny, Schliewen &amp; Dominey, 1992)</synonym>
	<synonym relationship='synonym'>Tilapia gutturosa Stiassny, Schliewen &amp; Dominey, 1992</synonym>
	<dcterms:created>1994-06-13</dcterms:created>
	<dcterms:modified>2024-04-18</dcterms:modified>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=6376&amp;speccode=10675'>IUCN0 1994 IUCN red list of threatened animals. International Union for Conservation of Nature and Natural Resources, Gland, Switzerland and Cambridge, U.K. (Ref. 6376)</reference>
	<reference isbn='0-86622-422-X.' url='http://www.fishbase.org/references/FBRefSummary.php?id=12251&amp;speccode=10675'>Axelrod, H.R.0 The most complete colored lexicon of cichlids. T.F.H. Publications, Neptune City, New Jersey. 864 p. (Ref. 12251)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=12255&amp;speccode=10675'>Baillie, J. and B. Groombridge (eds.)0 1996 IUCN red list of threatened animals. IUCN, Gland, Switzerland. 378 p. (Ref. 12255)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=12818&amp;speccode=10675'>Anonymous0 Fish registrations within the museum database of the Vertebrate Section of the Royal Museum for Central Africa. MRAC, Tervuren, Belgium. (Ref. 12818)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=26738&amp;speccode=10675'>Stiassny, M.L.J., U.K. Schliewen and W.J. Dominey0 A new species flock of cichlid fishes from Lake Bermin, Cameroon with a description of eight new species of &lt;i&gt;Tilapia&lt;/i&gt; (Labroidei: Cichlidae). Ichthyol. Explor. Freshwat. 3(4):311-346. (Ref. 26738)</reference>
	<reference isbn='957-8596-59-6' url='http://www.fishbase.org/references/FBRefSummary.php?id=31517&amp;speccode=10675'>Wu, H.L., K.-T. Shao and C.F. Lai (eds.)0 Latin-Chinese dictionary of fishes names. The Sueichan Press, Taiwan. 1028 p. (Ref. 31517)</reference>
	<reference isbn='2-8317-0564-9' url='http://www.fishbase.org/references/FBRefSummary.php?id=36508&amp;speccode=10675'>Hilton-Taylor, C.0 2000 IUCN red list of threatened species. IUCN, Gland, Switzerland and Cambridge, UK. xviii + 61 p. (with 1 CD-ROM). (Ref. 36508)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=41414&amp;speccode=10675'>Anonymous0 Fish collection database of the American Museum of Natural History. American Museum of Natural History, Central Park West , NY 10024-5192, USA. (Ref. 41414)</reference>
	<reference isbn='3-928819-33-X' url='http://www.fishbase.org/references/FBRefSummary.php?id=52307&amp;speccode=10675'>Lamboj, A.0 The cichlid fishes of Western Africa. Birgit Schmettkamp Verlag, Bornheim, Germany. 255 p. (Ref. 52307)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=58108&amp;speccode=10675'>Chinese Academy of Fishery Sciences0 Chinese aquatic germplasm resources database. http://zzzy.fishinfo.cn (Ref. 58108)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=81260&amp;speccode=10675'>Stiassny, M.L.J., A. Lamboj, D. De Weirdt and G.G. Teugels0 Cichlidae. p. 269-403. In M.L.J. Stiassny, G.G Teugels and C.D. Hopkins (eds.) The fresh and brackish water fishes of Lower Guinea, West-Central Africa Volume 2. Coll. faune et flore tropicales 42. Institut de recherche de développement, Paris, France, Muséum national d'histoire naturelle, Paris, France and Musée royal de l'Afrique Central, Tervuren, Belgium, 603p. (Ref. 81260)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=83882&amp;speccode=10675'>FAO-FIES0 Aquatic Sciences and Fisheries Information System (ASFIS) species list. Retrieved from http://www.fao.org/fishery/collection/asfis/en, March 2010. (Ref. 83882)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=90062&amp;speccode=10675'>FAO-FIES0 Aquatic Sciences and Fisheries Information System (ASFIS) species list. Retrieved from http://www.fao.org/fishery/collection/asfis/en, March 2012. (Ref. 90062)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=93285&amp;speccode=10675'>Dunz, A.R. and U.K. Schliewen0 Molecular phylogeny and revised classification of the haplotilapiine cichlid fishes formerly referred to as "&lt;i&gt;Tilapia&lt;/i&gt;". Mol. Phylogenet. Evol. 68(1):64-80. (Ref. 93285)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=95632&amp;speccode=10675'>FAO-FIES0 Aquatic Sciences and Fisheries Information System (ASFIS) species list. Retrieved from http://www.fao.org/fishery/collection/asfis/en, April 2014. (Ref. 95632)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=101110&amp;speccode=10675'>FAO-FIES0 Aquatic Sciences and Fisheries Information System (ASFIS) species list. Retrieved from http://www.fao.org/fishery/collection/asfis/en, [accessed 13/04/2015]. (Ref. 101110)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=123477&amp;speccode=10675'>ECoF, Eschmeyer's Catalog of Fishes0 Catalog of Fishes. 1985 - onwards. &lt;b&gt;CITATION NOT TO BE USED OUTSIDE FISHBASE.  SEE REMARKS&lt;/b&gt;. Authorship varies in time: Eschmeyer, W.N.; Fricke, R.; Van der Laan, R.  Web database. http://researcharchive.calacademy.org/research/ichthyology/catalog/fishcatmain.asp (Ref. 123477)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=130435&amp;speccode=10675'>IUCN0 The IUCN Red List of Threatened Species. 1964 - onwards. https://www.iucnredlist.org/ (Ref. 130435)</reference><dataObject>
	<dc:identifier>FB-GeneralDescr-10675</dc:identifier>
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	<agent homepage='' logoURL='' role='recorder'>Tess Cruz</agent>
	<dcterms:created>1994-06-13 00:00:00</dcterms:created>
	<dcterms:modified>2024-04-18 00:00:00</dcterms:modified>
	<dc:title>Biology</dc:title>
	<dc:language></dc:language>
	<license>http://creativecommons.org/licenses/by-nc/3.0/</license>
	<dc:rights></dc:rights>
	<dcterms:rightsHolder>WorldFish Center - FishBase</dcterms:rightsHolder>
	<dcterms:bibliographicCitation></dcterms:bibliographicCitation>
	<audience>General public</audience>
	<dc:source>http://www.fishbase.org/summary/SpeciesSummary.php?id=10675</dc:source>
	<subject>http://rs.tdwg.org/ontology/voc/SPMInfoItems#DiagnosticDescription</subject>
	<dc:description>Prefers sponges, but small amounts of diatoms, plants (mostly algae) and detritus have also been found in their stomachs (Ref. 52307). Pair-bonding, open (Ref. 52307) substrate brooder/spawner (Ref. 52307, 81260) with both parents guarding the brood (Ref. 52307). Usually spawns in shallow waters, building colonies of up to 6 pairs separated by about 30 cm; eggs are hidden between leaves and pieces of wood (Ref. 52307).</dc:description>
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	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=93285&amp;speccode=10675'>Dunz, A.R. and U.K. Schliewen 2013 Molecular phylogeny and revised classification of the haplotilapiine cichlid fishes formerly referred to as "&lt;i&gt;Tilapia&lt;/i&gt;". Mol. Phylogenet. Evol. 68(1):64-80. (Ref. 93285)</reference></dataObject><dataObject>
	<dc:identifier>FB-DiagnosticDescr-10675</dc:identifier>
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	<agent homepage='' logoURL='' role='recorder'>Armi G. Torres</agent>
	<dcterms:created>2007-01-08 00:00:00</dcterms:created>
	<dcterms:modified>2017-12-20 22:26:21</dcterms:modified>
	<dc:title></dc:title>
	<dc:language></dc:language>
	<license>http://creativecommons.org/licenses/by-nc/3.0/</license>
	<dc:rights></dc:rights>
	<dcterms:rightsHolder>WorldFish Center - FishBase</dcterms:rightsHolder>
	<dcterms:bibliographicCitation></dcterms:bibliographicCitation>
	<audience>General public</audience>
	<dc:source>http://www.fishbase.org/summary/SpeciesSummary.php?id=10675</dc:source>
	<subject>http://rs.tdwg.org/ontology/voc/SPMInfoItems#Morphology</subject>
	<dc:description>Diagnosis: lower pharyngeal jaw robust and massive, with a large, heart-shaped dentigerous plate, covered with numerous fine, regularly arrayed teeth; mouth not squared or obtuse; ventral margin of lower lip smooth; ratio of lower jaw length to lower jaw width &gt; 1; oral jaw teeth fixed with non-spatulate crowns (Ref. 26738, 81260). 13-16 gill rakers on first arch (Ref. 26738).Description: head length 34.4-38.5% SL; lower pharyngeal jaw with curved unicuspid anterior teeth, bicuspid posterior teeth; ventral keel shorter than dentigerous plate; 3 rows of teeth in upper jaw, outer row teeth bicuspid (Ref. 81260).Coloration: three colour forms are present in the lake and as far as can be determined they do not exhibit assortative mating. (1) "Normal" form: base body color greenish brown dorsally, blood-red ventrally; snout and dorsum of head greenish brown (Ref. 81260). Upper lip black, lower lip pale (Ref. 52307, 81260) with turquoise iridescence (Ref. 81260); cheek and opercle blood-red ventrally, becoming yellowish green dorsally; chest bright blood-red; 5-6 faint vertical bars often present on flanks (Ref. 81260). Dorsal fin yellowish brown; anal fin red with dusky black distal margin; caudal fin yellow-brown with dusky black distal margin and red ventral lobe (Ref. 52307, 81260). Pelvics smoky red; pectoral base yellow, fin smoky brown (Ref. 81260). (2) "Bronze" form: base body color bronze-brown dorsally, yellowish bronze ventrally; snout and dorsum of head brownish black; upper lip black, lower lip pale with turquoise iridescence; cheek and opercle reddish bronze; chest yellow-bronze; 5-6 faint vertical bars often present on flanks; dorsal fin brownish bronze; caudal fin brown with smoky distal margin; anal fin brown with smoky black distal margin; pelvics smoky grey; pectoral base yellow, fin pale smoky yellow; this is a common female coloration, but individuals exhibiting "normal" or "pale" coloration are known to occur (Ref. 81260). (3) "Pale" form: base body color pale brown dorsally, silvery grey ventrally (Ref. 52307, 81260). Snout and upper lip smoky grey, lower lip silvery white; throat, chest and belly silvery white; dorsal fin pale brownish yellow with red tinge to lappets; prominent "tilapia spot" usually present; caudal fin margin with reddish tinge; anal, pelvic and pectoral fins transparent (Ref. 81260).</dc:description>
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	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=81260&amp;speccode=10675'>Stiassny, M.L.J., A. Lamboj, D. De Weirdt and G.G. Teugels 2008 Cichlidae. p. 269-403. In M.L.J. Stiassny, G.G Teugels and C.D. Hopkins (eds.) The fresh and brackish water fishes of Lower Guinea, West-Central Africa Volume 2. Coll. faune et flore tropicales 42. Institut de recherche de développement, Paris, France, Muséum national d'histoire naturelle, Paris, France and Musée royal de l'Afrique Central, Tervuren, Belgium, 603p. (Ref. 81260)</reference></dataObject><dataObject>
	<dc:identifier>FB-Morphology-10675</dc:identifier>
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	<agent homepage='' logoURL='' role='recorder'>Armi G. Torres</agent>
	<dcterms:created>2007-01-08 00:00:00</dcterms:created>
	<dcterms:modified>2017-12-20 22:26:21</dcterms:modified>
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	<dc:language></dc:language>
	<license>http://creativecommons.org/licenses/by-nc/3.0/</license>
	<dc:rights></dc:rights>
	<dcterms:rightsHolder>WorldFish Center - FishBase</dcterms:rightsHolder>
	<dcterms:bibliographicCitation></dcterms:bibliographicCitation>
	<audience>General public</audience>
	<dc:source>http://www.fishbase.org/summary/SpeciesSummary.php?id=10675</dc:source>
	<subject>http://rs.tdwg.org/ontology/voc/SPMInfoItems#Distribution</subject>
	<dc:description>Dorsal spines (total): 14 - 16; Dorsal soft rays (total): 10 - 11;  Analspines: 3; Analsoft rays: 7 - 8</dc:description>
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	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=81260&amp;speccode=10675'>Stiassny, M.L.J., A. Lamboj, D. De Weirdt and G.G. Teugels 2008 Cichlidae. p. 269-403. In M.L.J. Stiassny, G.G Teugels and C.D. Hopkins (eds.) The fresh and brackish water fishes of Lower Guinea, West-Central Africa Volume 2. Coll. faune et flore tropicales 42. Institut de recherche de développement, Paris, France, Muséum national d'histoire naturelle, Paris, France and Musée royal de l'Afrique Central, Tervuren, Belgium, 603p. (Ref. 81260)</reference></dataObject><dataObject>
	<dc:identifier>FB-Distribution-10675</dc:identifier>
	<dataType>http://purl.org/dc/dcmitype/Text</dataType>
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	<agent homepage='' logoURL='' role='recorder'>Tess Cruz</agent>
	<dcterms:created>1994-06-13 00:00:00</dcterms:created>
	<dcterms:modified>2025-11-05 00:00:00</dcterms:modified>
	<dc:title></dc:title>
	<dc:language></dc:language>
	<license>http://creativecommons.org/licenses/by-nc/3.0/</license>
	<dc:rights></dc:rights>
	<dcterms:rightsHolder>WorldFish Center - FishBase</dcterms:rightsHolder>
	<dcterms:bibliographicCitation></dcterms:bibliographicCitation>
	<audience>General public</audience>
	<dc:source>http://www.fishbase.org/summary/SpeciesSummary.php?id=10675</dc:source>
	<subject>http://rs.tdwg.org/ontology/voc/SPMInfoItems#Uses</subject>
	<dc:description>Africa:  endemic to Lake Bermin, Cameroon (Ref. 52307, 81260).</dc:description>
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	<location></location>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=130435&amp;speccode=10675'>IUCN 2025 The IUCN Red List of Threatened Species. 1964 - onwards. https://www.iucnredlist.org/ (Ref. 130435)</reference></dataObject><dataObject>
	<dc:identifier>FB-Size-10675</dc:identifier>
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	<agent homepage='' logoURL='' role='recorder'>Tess Cruz</agent>
	<dcterms:created>1994-06-13 00:00:00</dcterms:created>
	<dcterms:modified>2024-04-18 00:00:00</dcterms:modified>
	<dc:title>Max. size</dc:title>
	<dc:language></dc:language>
	<license>http://creativecommons.org/licenses/by-nc/3.0/</license>
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	<dcterms:bibliographicCitation></dcterms:bibliographicCitation>
	<dc:source>http://www.fishbase.org/summary/SpeciesSummary.php?id=10675</dc:source>
	<dc:description>6.4 cm SL (male/unsexed; (Ref. 12251))</dc:description>
	<mediaURL></mediaURL>
	<thumbnailURL></thumbnailURL>
	<location></location>
	<reference isbn='0-86622-422-X.' url='http://www.fishbase.org/references/FBRefSummary.php?id=12251&amp;speccode=10675'>Axelrod, H.R. 1993 The most complete colored lexicon of cichlids. T.F.H. Publications, Neptune City, New Jersey. 864 p. (Ref. 12251)</reference></dataObject><dataObject>
	<dc:identifier>FB-Threats-2</dc:identifier>
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	<agent homepage='' logoURL='' role='recorder'>Belen Acosta</agent>
	<dcterms:created>1990-10-18 00:00:00</dcterms:created>
	<dcterms:modified>2024-03-07 00:00:00</dcterms:modified>
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	<audience>General public</audience>
	<dc:source>http://www.fishbase.org/summary/SpeciesSummary.php?id=2</dc:source>
	<subject>http://rs.tdwg.org/ontology/voc/SPMInfoItems#Size</subject>
	<dc:description>Least Concern (LC) , IUCN Grouper and Wrasse Specialist Group</dc:description>
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	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=57073'>IUCN 2006 2006 IUCN red list of threatened species. www.iucnredlist.org. Downloaded July 2006.</reference></dataObject><dataObject>
	<dc:identifier>FB-Habitat-10675</dc:identifier>
	<dataType>http://purl.org/dc/dcmitype/Text</dataType>
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	<agent homepage='' logoURL='' role='recorder'>Tess Cruz</agent>
	<dcterms:created>1994-06-13 00:00:00</dcterms:created>
	<dcterms:modified>2024-04-18 00:00:00</dcterms:modified>
	<dc:title>Environment</dc:title>
	<dc:language></dc:language>
	<license>http://creativecommons.org/licenses/by-nc/3.0/</license>
	<dc:rights></dc:rights>
	<dcterms:rightsHolder>WorldFish Center - FishBase</dcterms:rightsHolder>
	<dcterms:bibliographicCitation></dcterms:bibliographicCitation>
	<audience>General public</audience>
	<dc:source>http://www.fishbase.org/summary/SpeciesSummary.php?id=10675</dc:source>
	<subject>http://rs.tdwg.org/ontology/voc/SPMInfoItems#Threats</subject>
	<dc:description>benthopelagic; freshwater</dc:description>
	<mediaURL></mediaURL>
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