Females: esca without distal prolongation of bulb; a single distal branched filament, length less than half diameter of bulb; three branched filaments along each side of bulb, longest 1-3 times diameter of bulb; no posterior appendage. Barbel 24-80% SL in total length; distal half to two third divided into three main-branches, proximal, posterior main-branch simple, anterior main-branch distally divided into 4 filaments, median main branch bifurcated; each secondary branch with one or more filaments; all filaments unpigmented, some branched, each with a series of sessile photophores. Dense concentration of subdermal pigment on caudal peduncle (Ref. 42949). Males: with pointed sphenotic spines (Ref. 86949).