Diagnosis: Distinct from its congeners in having a round or trapezoidal black humeral spot, dorsal to the lateral line; a branch of the infraorbital sensory canal directed ventrally, continued as a series of pores totally or partially traversing the third infraorbital; body relatively slender (31.8-37.4% SL); eye large (41.1-45.8% HL); snout short (15.5-20.9% HL); caudal peduncle relatively slender (10.3-12% SL); one pentacuspid maxillary tooth; pored lateral line scales 38-42; and anal fin rays v-vi, 23-29. The males have breeding tubercles on the head and scales, and bony hooks on all fins (Ref. 57546).