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| Sound produced | yes, active sound production |
| Type de son produit | thumps |
| Organe sonore | swim bladder |
| Sound mechanism | vibration by contraction of associated muscles and general body contraction |
| Contexte comportemental | under duress (handling or cornering) |
| Reference | Fish, M.P. and W.H. Mowbray, 1970 |
| Remarque |