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| Sound produced | yes, active sound production |
| 発生した音の種類 | knocks, single or pulsed drum beats |
| 音響発生器官 | swim bladder |
| Sound mechanism | vibration by contraction of primary sonic muscles & conformation of swim bladder |
| 行動状況 | intraspecific competition, terretorial behaviour |
| Reference | Parmentier, E., P. Vandewalle and J.P. Lagardère, 2003 |
| 意見 | strongly species-specific sounds, aquarium recording, recorded with lowpass filter: 6400 Hz. |