Africa: middle Congo River basin in the Kwilu (Ref. 11970), Kwango, Kasai drainage (Ref. 11970, 55074) and Lomami (Ref. 106245); upper Congo River basin including the Lualaba, upper Lualaba, Lufira, Upemba region, Luvua, Lake Mweru, Luapula (Ref. 55074) and Bangweulu (Ref. 5163, 95585); Cunene, Okavango, Lake Ngami, Zambezi, Limpopo, northern tributaries of the Orange River (Ref. 5163, 11970), Cuvelai (Ref. 120641), upper Cuanza, Sabi, Lundi and Lake Malawi (Ref. 5163).