South America: Magdalena River basin; entire extent of the Amazon and Orinoco River basins below approximately 500 m elevation including Napo River of western Ecuador and Madre de Dios River of Peru; all drainages of the Guianas Shield; Atlantic drainages of northeastern Brazil (Tocantins, Parnaíba, Pindaré, Itapicuru, and Salgado rivers); São Francisco River basin; Rio de Janeiro, Brazil; Paraguay-Paraná River basin.