| اقتباس | Ross, S.T., 1973, The systematics of Gasterosteus aculeatus (Pisces, Gasterosteidae) in central and southern Califormia.. Natur. Hist. Mus. Los Angeles Country. Contrib. Sci. 243:1-20. |
| مرجع | 35064 Ross, S.T., 1973 |
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| Named Used as Valid | Gasterosteus aculeatus williamsoni |
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