| Citation | Klinkhardt, M, M. Tesche and H. Greven, 1995, Database of fish chromosomes.. Westarp Wissenschaften. |
| Ref. | 34370 Klinkhardt, M, M. Tesche and H. Greven, 1995 |
| Page | 132 |
| Named Used as Valid | Gasterosteus aculeatus williamsoni |
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