| Citation | Severin, S.O., 1986, Divergence of graylings (the genus Thymallus Cuvier, 1829) on the basis of karyological data.. Zool. Zh . 65(8):1190-1198. |
| Ref. | 29978 Severin, S.O., 1986 |
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| Named Used as Valid | Thymallus thymallus |
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